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Type: Article
Published: 2008-09-19
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Phytoseiid mites (Acari: Phytoseiidae) of the subfamily Amblyseiinae Muma from Peru, with descriptions of four new species

Depto. Ent. Fitop. e Zool. Agrícola, ESALQ/Universidade de São Paulo, 13418-900 Piracicaba – SP, Brazil
Depto. Ent. Fitop. e Zool. Agrícola, ESALQ/Universidade de São Paulo, 13418-900 Piracicaba – SP, Brazil
International Centre of Insect Physiology and Ecology, P.O. Box 30772, 00100 Nairobi, Kenya, current address: Koppert Biological Systems, Postbus 155, 2650 AD Berkel en Rodenrijs, The Netherlands
Acari Taxonomy predatory mites biological control

Abstract

This paper reports the mites of the subfamily Amblyseiinae (Phytoseiidae) known from Peru, based on an extensive survey conducted in different regions of the country as well as on published information. Forty-eight species are reported, four of which are new to science, namely Aristadromips escarsidentatus Guanilo & Moraes n. sp., Proprioseiopsis parabelizensis Guanilo & Moraes n. sp., Typhlodromips angustus Guanilo & Moraes n. sp. and Typhlodromips amazonensis Guanilo & Moraes n. sp. In addition to the description of the new species, measurements of all species collected and a taxonomic key to separate the species reported from Peru are provided.

References

  1. Aponte, O. & McMurtry, J.A. (1995) Revision of the genus Iphiseiodes De Leon (Acari: Phytoseiidae). International Journal of Acarology, 21, 165–183.

    Athias-Henriot, C. (1957) Phytoseiidae et Aceosejidae (Acarina, Gamasina) d'Algerie. I Genres Blattisocius Keegan, Iphiseius Berlese, Amblyseius Berlese, Phytoseius Ribaga, Phytoseiulus Evans. Bulletin de la Societe d’Histoire Naturelle de l’Afrique du Nord, 48, 319–352.

    Athias-Henriot, C. (1967) Nouveaux Amblyseius edaphiques d' Amerique Australe (Acariens anactinotriches, Phytoseiidae). In: Deboutteville, C.D. & Rapoport, E. (Eds.), Biologie de l'Amerique Australe. III. Etudes sur la Faune du Sol. Documents Biogeographiques. Centre National Recherche Scientifique, Paris, France, pp. 525–539.

    Athias-Henriot, C. (1977) Nouvelles notes sur les Amblyseiini. III. Sur le genre Cydnodromus: Redefinition, composition (Parasitiformes, Phytoseiidae). Entomophaga, 22, 61–73.

    Banks, N. (1905) Descriptions of some new mites. Proceedings of the Entomological Society of Washington, 7, 133–142.

    Berlese, A. (1914) Acari nuovi. Manipulus IX. Redia, 10, 113–150.

    Chant, D.A. (1957) Note on the status of some genera in the family Phytoseiidae (Acarina). The Canadian Entomologist, Canada, 89(11), 528–532.

    Chant, D.A. (1959) Phytoseiid mites (Acarina: Phytoseiidae). Part I. Bionomics of seven species in southeastern England. Part II. A taxonomic review of the family Phytoseiidae, with descriptions of thirty-eight new species. The Canadian Entomologist, Supplement 12, 1–166.

    Chant, D.A. & Baker, E.W. (1965) The Phytoseiidae (Acarina) of Central America. Memoirs of the Entomological Society of Canada, 41, 1–56.

    Chant, D.A. & McMurtry, J.A. (2003) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part I. Neoseiulini new tribe. International Journal of Acarology, 29, 3–46.

    Chant, D.A. & McMurtry, J.A. (2004a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part III. The tribe Amblyseiini Wainstein. International Journal of Acarology, 30, 171–228.

    Chant, D. A. & McMurtry, J. A. (2004b) A review of the subfamily Amblyseiini Muma (Acari: Phytoseiidae). Part IV. The tribe Amblyseiinae Muma Wainstein, subtribe Arrenoseiina Chant & McMurtry. International Journal of Acarology. 30, 291–312.

    Chant, D.A. & McMurtry, J.A. (2005a) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part V. Tribe Amblyseiini, subtribe Proprioseiopsina Chant & McMurtry. International Journal of Acarology, 31, 3–22.

    Chant, D.A. & McMurtry, J.A. (2005b) A review of the subfamily Amblyseiinae Muma (Acari: Phytoseiidae): Part VI. The tribe Euseiini n. tribe, subtribes Typhlodromalina n. subtribe, Euseiina n. subtribe, and Ricoseiina n. subtribe. International Journal of Acarology, 31, 187–224.

    Chant, D.A. & McMurtry, J.A. (2005c) A review of the subfamily Amblyseiinae Muma. (Acari: Phytoseiidae): Part VII. Typhlodromipsini n. tribe. International Journal of Acarology, 31, 315–340.

    Chant, D.A. & McMurtry, J.A. (2006) A review of the subfamily Amblyseiinae Muma. (Acari: Phytoseiidae): Part VIII. The tribes Macroseiini Chant, Denmark and Baker, Phytoseiulini N. Tribe, Afroseiulini N. Tribe and Indoseiulini Ehara and Amano. International Journal of Acarology, 32, 13 – 25.

    Chant, D.A. & McMurtry, J.A. (2007) Illustrated keys and diagnoses for the genera and subgenera of the Phytoseiidae of the world (Acari: Mesostigmata). Indira Publishing House; West Bloomfield, USA. pp 219.

    Chant, D.A. & Yoshida-Shaul, E. (1991) Adult ventral setal patterns in the family Phytoseiidae (Acari: Gamasina). International Journal of Acarology, 17, 187–199.

    Chant, D.A., Denmark, H.A. & Baker, E.W. (1959) A new subfamily, Macroseiinae Nov., of the family Phytoseiidae (Acarina: Gamasina). The Canadian Entomologist, 91, 808–811.

    Chant, D.A., Hansell, R.I.C., Rowell, H.J. & Yoshida-Shaul E. (1978) A study of the family Phytoseiidae (Acarina: Mesostigmata) using the methods of numerical taxonomy. Canadian Journal of Zoology, 56, 1330–1347.

    Chaudhri, W.M. (1968) Six new species of mites of the genus Amblyseius (Phytoseiidae) from Pakistan. Acarologia, 10, 550–562.

    De Leon, D. (1961) Eight new Amblyseius from Mexico with collection notes on two other species (Acarina: Phytoseiidae). The Florida Entomologist, 44, 85–91.

    De Leon, D. (1962) The cervices of some phytoseiid type specimens (Acarina: Phytoseiidae). Acarologia, 4, 174–176.

    De Leon, D. (1965a) Phytoseiid mites from Puerto Rico with descriptions of new species (Acarina: Mesostigmata). The Florida Entomologist, 48(2), 121–131.

    De Leon, D. (1965b) A note on Neoseiulus Hughes 1948 and new synonymy (Acarina: Phytoseiidae). Proceedings of the Entomological Society of Washington, 67, 23.

    De Leon, D. (1966) Phytoseiidae of British Guyana with keys to species (Acarina: Mesostigmata). Studies on the Fauna of Suriname and other Guyanas, 8, 81–102.

    De Leon, D. (1967) Some mites of the Caribbean Area. Part I. Acarina on plants in Trinidad, West Indies. Allen Press Inc., Lawrence, 66 pp.

    Daneshvar, H. & Denmark, H.A. (1982) Phytoseiids of Iran (Acarina: Phytoseiidae). International Journal of Acarology, 8, 3–14.

    Denmark, H.A. (1965) Four new Phytoseiidae (Acari: Mesostigmata) from Florida. The Florida Entomologist, 48, 89–95.

    Denmark, H.A. & Muma, M.H. (1967) Six new Phytoseiidae from Florida (Acarina: Phytoseiidae). The Florida Entomologist, 50, 169–180.

    Denmark, H.A. & Muma, M.H. (1970) Some phytoseiid mites of Paraguay (Phytoseiidae: Acarina). The Florida Entomologist, 53, 219–227.

    Denmark, H.A. & Muma, M.H. (1972) Some Phytoseiidae of Colombia (Acarina: Phytoseiidae). The Florida Entomologist, 55, 19–29.

    Denmark, H.A. & Muma, M.H. (1973) Phytoseiid mites of Brazil (Acarina: Phytoseiidae). Revista Brasileira de Biologia, 33, 235–276.

    Denmark, H.A. & Muma, M.H. (1975) The Phytoseiidae (Acarina: Mesostigmata) of Puerto Rico. The Journal of Agriculture of the University of Puerto Rico, 59, 279–304.

    Denmark, H.A. & Muma, M.H. (1989) A revision of the genus Amblyseius Berlese, 1914 (Acari: Phytoseiidae). Occasional Papers of the Florida State Collection of Arthropods, 4, 1–149.

    Denmark, H.A. & Schicha, E. (1983) Revision of the genus Phytoseiulus Evans (Acarina: Phytoseiidae). International Journal of Acarology, 9, 27–35.

    Denmark, H.A., Evans, G.A., Aguilar, H., Vargas, C. & Ochoa, R. (1999) Phytoseiidae of Central America (Acari: Mesostigmata). Indira Publishing House, West Bloomfield, 125 pp.

    Dosse, G. (1958) Uber einige neue Raubmilbenarten (Acari: Phytoseiidae). Pflanzenschutz Berichte, 21, 44–61.

    Ehara, S. (1966) Some mites associated with plants in the state of Sao Paulo, Brazil, with a list of plant mites of South America. Japanese Journal of Zoology, 15, 129–150.

    Ehara, S. & Amano, H. (1998) A revision of the mite family Phytoseiidae in Japan (Acari: Gamasina), with remarks on its biology. Species Diversity, 3, 25–73.

    El-Banhawy, E.M. (1979) Records on phytoseiid (Acari) mites of Peru. International Journal of Acarology, 5, 111–116.

    El-Banhawy, E.M. (1984) Description of some phytoseiid mites from Brazil (Acarina: Phytoseiidae). Acarologia, 25, 125–144.

    Evans, G.O. (1952) On a new predatory mite of economic importance. Bulletin of Entomological Research, 43, 397–401.

    Feres, R.J.F. (2000) Levantamento e observações naturalísticas da acarofauna (Acari, Arachnida) de seringueiras cultivadas (Hevea spp., Euphorbiaceae) no Brasil. Revista Brasileira de Zoologia, 17, 157–173.

    Feres, R.J.F. & Moraes G.J. de, (1998) Phytoseiid mites (Acari: Phytoseiidae) from woody areas in the State of Paulo, Brazil. Systematic and Applied Acarology, 3, 125–132.

    Feres, R.J.F. & Nunes, M.A. (2001) Ácaros (Acari, Arachnida) associados a euforbiáceas nativas em áreas de cultivo de seringueiras (Hevea brasiliensis Muell. Arg., Euphorbiaceae) na região noroeste do Estado de São Paulo, Brasil. Revista Brasileira de Zoologia, 18, 1253–1264.

    Feres, R.J.F., Rossa-Feres, D. de C., Daud, R.D. & Santos, R.S. (2002) Diversidade de ácaros (Acari, Arachnida) em seringueiras (Hevea brasiliensis Muell. Arg., Euphorbiaceae) na região noroeste do Estado de São Paulo, Brasil. Revista Brasileira de Zoologia, 19, 137–144.

    Ferla, N.J. & Moraes, G.J. de (2002) Ácaros predadores (Acari) em plantas nativas e cultivadas do Estado do Rio Grande do Sul, Brasil. Revista Brasileira de Zoologia, 19, 1011–1031.

    Fiaboe, K.K.M., Fonseca, R.L., Moraes, G.J. de, Ogol, C.K.P.O. & Knapp, M. (2006) Identification of priority areas in South America for exploration of natural enemies for classical biological control of Tetranychus evansi (Acari: Tetranychidae) in Africa. Biological Control, 38, 373–379.

    Fiaboe, K.K.M., Gondim Jr, M.G.C., Moraes, G.J. de, Ogol, C.K.P.O. & Knapp, M. (2007) Surveys for natural enemies of the tomato red spider mite Tetranychus evansi (Acari: Tetranychidae) in the northeastern and southeastern Brazil. Zootaxa, 1395, 33–58.

    Furtado, I.P., Kreiter, S., Moraes, G.J. de, Tixier, M.S., Flechtmann, C.H.W. & Knapp, M. (2005) Plant mites (Acari) from northeastern Brazil, with descriptions of two new species of the family Phytoseiidae (Mesostigmata). Acarologia, 45, 131–143.

    Furtado, I.P., Moraes, G.J. de, Kreiter, S. & Knapp, M. (2006) Search for effective natural enemies of Tetranychus evansi in south and southeast Brazil. Experimental and Applied Acarology, 40, 157–174.

    Furtado, I.P., Toledo, S. E., Moraes, G.J. de, Kreiter, E.S. & Knapp, M. (2007) Search for effective natural enemies of Tetranychus evansi (Acari: Tetranychidae) in northwest Argentina. Experimental and Applied Acarology, 43, 121–127.

    Garman, P. (1958) New species belonging to the genera Amblyseius and Amblyseiopsis with keys to Amblyseius, Amblyseiopsis and Phytoseiulus. Annals of the Entomological Society of America, 51, 69–79.

    Gondim Jr, M.G.C. & Moraes, G.J. de (2001) Phytoseiid mites (Acari: Phytoseiidae) associated with palm trees (Arecaceae) in Brazil. Systematic and Applied Acarology, 6, 65–94.

    Gonzalez, R.H. & Schuster, R.O. (1962) Especies de la familia Phytoseiidae en Chile I. (Acarina: Mesostigmata). Boletín Técnico. Estación Experimental Agronómica. Universidad de Chile, Facultad de Agronomía, Chile, 16, 1–35.

    Guanilo, A. D. & Martínez, N. (2007) Predators associated to Panonychus citri McGregor (Acari: Tetranychidae) in the Central Coast of Peru. Ecologia Aplicada, Peru, 5, 117–129 [in Spanish].

    Hirschmann W. (1962) Gangystematik der Parasitiformes. Acarologie Schriftenreihe fur Vergleichende Milbenkunde, 5, 1–80 pp. + 32 plates.

    Hughes, A.M. (1948) The mites associated with stored food products. Ministry of Agriculture and Fishery, H. M. Stationery Office, London, 168 pp.

    Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes): Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1–475.

    Karg, W. (1983) Systematische untersuchung der Gattungen und Untergattungen der Raubmilbenfamilie Phytoseiidae Berlese, 1916, mit der beschreibung von 8 neuen Arten. Mitteilungen Zoologisches Museum in Berlin, 59, 293–328.

    Karg, W. (1989) Neue Raubmilbenarten der Gattung Proprioseiopsis Muma, 1961 (Acarina, Parasitiformes) mit Bestimmungsschlusseln. Zoologische Jahrbucher Systematik, 116, 199–216.

    Kennett, C.E. (1958) Some predacious mites of the subfamilies Phytoseiinae and Aceosejinae (Acarina: Phytoseiidae, Aceosejidae) from central California with descriptions of new species. Annals of the Entomological Society of America, 51, 471–479.

    Kennett, C.E. & Caltagirone, L.E. (1968) Biosystematics of Phytoseiulus persimilis Athias-Henriot (Acarina: Phytoseiidae). Acarologia, 10, 563–577.

    Kreiter, S. & Moraes, G.J. de (1997) Phytoseiidae mites (Acari: Phytoseiidae) from Guadeloupe and Martinique. The Florida Entomologist, 80, 376–382.

    Lofego, A.C., Moraes, G.J. de & Castro, L.A.S. (2004) Phytoseiid mites (Acari: Phytoseiidae) on Myrtaceae in the State of São Paulo, Brazil. Zootaxa. 516, 1–18.

    Lombardini, G. (1959) Acari Nuovi. XXXVII. Bollettino dell’Istituto di Entomologia Agraria, della Universita di Palermo ed Osservatorio Regionale per le Malattie delle Piante, 21, 163–167.

    McGregor, E.A. (1954) Two new mites in the genus Typhlodromus (Acarina: Phytoseiidae). Southern California Academy of Science Bulletin, 53, 89–92.

    McMurtry, J.A. (1983) Phytoseiid mites from Guatemala, with descriptions of two new species and redefinitions of the genera Euseius, Typhloseiopsis, and the Typhlodromus occidentalis species group (Acari: Mesostigmata). International Journal of Entomology, 25, 249–272.

    McMurtry, J.A. & Moraes, G.J.de (1984) Some phytoseiid mites from the South Pacific, with descriptions of new species and a definition of the Amblyseius largoensis species group. International Journal of Acarology, 10, 27–37.

    McMurtry, J.A. & Moraes, G.J.de (1989) Some phytoseiid mites from Peru with descriptions of four new species (Acari: Phytoseiidae). International Journal of Acarology, 15, 179–188.

    Moraes, G.J. de & McMurtry, J.A. (1983) Phytoseiid mites (Acarina) of northeastern Brazil with descriptions of four new species. International Journal of Acarology, 9, 131–148.

    Moraes, G.J. de & Mesa, N.C. (1988) Mites of the family Phytoseiidae (Acari) in Colombia, with descriptions of three new species. International Journal of Acarology, 14, 71– 88.

    Moraes, G.J. de, Denmark, H.A. & Guerrero, J.M. (1982) Phytoseiid mites of Colombia (Acarina: Phytoseiidae). International Journal of Acarology, 8, 15– 22.

    Moraes, G.J. de, Kreiter, S. & Lofego, A.C. (2000) Plant mites (Acari) of the French Antilles. 3. Phytoseiidae (Gamasida). Acarologia, 40, 237–264.

    Moraes, G.J. de; McMurtry, J.A. & Baker, E.W. (1987) Redescription and distribution of the spider mites Tetranychus evansi and T. marianae. Acarology, 28, 333–343.

    Moraes, G.J. de, McMurtry, J.A. & Denmark, H.A. (1986) A Aatalog of the Mite Family Phytoseiidae. References to Taxonomy, Synonymy, Distribution and Habitat. EMBRAPA - DDT, Brasilia, Brazil, 353 pp.

    Moraes, G.J. de, Mesa, N.C. & Braun, A. (1991) Some phytoseiid mites of Latin America (Acari: Phytoseiidae). International Journal of Acarology, 17, 117–139.

    Moraes, G.J. de, Mesa, N.C., Braun, A. & Melo, E.L. (1994) Definition of the Amblyseius limonicus species group (Acari: Phytoseiidae), with descriptions of two new species and new records. International Journal of Acarology, 20, 209–217.

    Moraes, G.J.de, McMurtry, J.A., Denmark, H.A. & Campos, C.B. (2004) A revised catalog of the mite family Phytoseiidae. Zootaxa, 434, 1–494.

    Moraes, G.J. de, Alencar, J.A. de, Lima, J.L.S., Yaninek, J.S. & Delalibera Jr., I. (1993) Alternative plant habitats for common phytoseiid predators of the cassava green mite (Acari: Phytoseiidae, Tetranychidae) in northeast Brazil. Experimental and Applied Acarology, 17, 77–90.

    Moraes, G. J., Zannou, I. D., Ueckermann, E., Oliveira, A. R., Hanna, R. & Yaninek, J. S. (2007) Species of the subtribes Arrenoseiina and Proprioseiopsina (tribe Amblyseiini) and the tribe Typhlodromipsini (Acari: Phytoseiidae) from sub-Saharan Africa. Zootaxa, 1448, 1–39.

    Muma, M.H. (1955) Phytoseiidae (Acarina) associated with citrus in Florida. Annals of the Entomological Society of America, 48, 262–272.

    Muma, M.H. (1961) Subfamiles, genera, and species of Phytoseiidae (Acarina: Mesostigmata). Florida State Museum Bulletin, 5, 267–302.

    Muma, M.H. (1964) The population of Phytoseiidae on Florida citrus. The Florida Entomologist, 7, 5–11.

    Muma, M.H., Denmark, H.A. & De Leon, D. (1970) Phytoseiidae of Florida. Arthropods of Florida and neighboring land areas, 6. Florida Department of Agriculture and Consumer Services, Division of Plant Industry, Gainesville, USA, 150 pp.

    Murphy, S.T. (1984) Biological control of the cassava green mite (Mononychellus spp.) in east Africa. Integrated Pest Management of Cassava Green Mite. Proceedings Regional Training Workshop in East Africa, 30 April - 4 May 1984. CIBC, Kenya, pp. 55–61.

    Noronha, A.C.S. & Moraes, G.J. de 2002. Variações morfológicas intra e interpopulacionais de Euseius citrifolius Denmark & Muma e Euseius concordis (Chant) (Acari, Phytoseiidae). Revista. Brasileira. de Zoologia. 19, 1111–1122.

    Nesbitt, H.H.J. (1951) A taxonomic study of the Phytoseiidae (Family Laelapidae) predaceous upon Tetranychidae of economic importance. Zoologische Verhandelingen, 12, 64 pp.+ 32 plates.

    Pritchard, A.E. & Baker, E.W. (1962) Mites of the family Phytoseiidae from Central Africa, with remarks on genera of the world. Hilgardia, 33, 205–309.

    Rosa, A.A., Gondim Jr., M.G.C., Fiaboe, K.K.M., Moraes, G.J. de & Knapp, M. (2005) Predatory mites associated with Tetranychus evansi Baker & Pritchard (Acari: Tetranychidae) on native solanaceous plants of coastal Pernambuco State, Brazil. Neotropical Entomology, 34, 689–692.

    Rowell, H.J., Chant, D.A. & Hansell, R.I.C. (1978) The determination of setal homologies and setal patterns on the dorsal shield in the family Phytoseiidae (Acarina: Mesostigmata). The Canadian Entomologist, 110, 859–876.

    Schicha, E. (1987) Phytoseiidae of Australia and Neighboring Areas. Indira Publishing House, West Bloomfield, Michigan, USA, 187 pp.

    Schicha, E. & Elshafie, M. (1980) Four new species of phytoseiid mites from Australia, and three species from America redescribed (Acari: Phytoseiidae). Journal of the Australian Entomological Society, 19, 27–36.

    Schuster, R.O. (1966) Phytoseiidae of the Galapagos Islands (Acarina: Mesostigmata). Pacific Insects, 8, 319–339.

    Schuster, R.O. & Pritchard, A.E. (1963) Phytoseiid mites of California. Hilgardia, 34, 191–285.

    Takahashi, F. & Chant, D.A. (1993) Phylogenetic relationships in the genus Phytoseiulus Evans (Acari: Phytoseiidae). II. Taxonomic review. International Journal of Acarology, 19, 23–37.

    Tseng, Y.H. (1983) Further study on phytoseiid mites from Taiwan (Acarina: Mesostigmata). Chinese Journal of Entomology, 3, 33–74.

    Van der Merwe, G.G. (1968) A taxonomic study of the family Phytoseiidae (Acari) in South Africa with contributions to the biology of two species. Entomology Memoirs South Africa Department of Agricultural Technical Services, 18, 1–198.

    Wainstein, B.A. (1962) Revision du genre Typhlodromus Scheuten, 1857 et systematique de la famille des Phytoseiidae (Berlese 1916) (Acarina: Parasitiformes). Acarologia, 4, 5–30.

    Wainstein, B.A. (1983) Predaceous mites of the family Phytoseiidae (Parasitiformes) of Hawaii. Entomological Review, 62, 181–186.

    Wu, W.N. & Chou, F.W. (1981) A new species of Amblyseius (Acarina: Phytoseiidae) from Guangdong Province [in Chinese]. Zoological Research, 2, 273–274.

    Yaninek, J.S, Onzo, A & Ojo, J. B. (1993) Continent-wide releases of neotropical phytoseiids against the exotic cassava green mite in Africa. Experimental and Applied Acarology, 17, 145–160.

    Yaseen, M. & Bennett, F.D. (1976) Distribution, biology and population dynamics of the green cassava mite in the neotropics. Proceedings of the Fourth Symposium of the International Society of Tropical Root Crops, CIAT, Cali, Colombia, 1 7 Aug. 1976, Colombia, pp. 197–202.

    Yoshida-Shaul, E. & Chant, D.A. (1988) Descriptions of two unusual new species in the genus Amblyseius Berlese (Acari: Phytoseiidae). Canadian Journal of Zoology, 66, 2053–2056.

    Yoshida-Shaul, E. & D. A. Chant (1991) Five new species of Phytoseiidae from Central and South America (Acari: Gamasina). International Journal of Acarology, 17, 93–102.

    Zacarias, M.S. & Moraes, G.J. de (2001) Phytoseiid mites (Acari) associated with rubber trees and other euphorbiaceous plants in southeastern Brazil. Neotropical Entomology, 30, 579–586.

    Zannou, I. D., Moraes, G. J. de; Ueckermann, E., Oliveira, A. R., Yaninek, J. S. & Hanna, R. (2006) Phytoseiid mites of the genus Neoseiulus Hughes (Acari: Phytoseiidae) from sub-Saharan Africa. International Journal of Acarology, 32, 241–276.