Abstract
Cyrtodactylus ayunpaensis sp. nov. from Ayun Pa town, Gia Lai Province, Central Highlands Vietnam is described based on an integrative taxonomic method by a combination of morphological and genetic data. It is placed into the C. irregularis group and can be distinguished from the remaining species from Vietnam by a minimum genetic distance of 12.8% in the ND2 gene and by the following characters: a maximum SVL of 91.5 mm; supralabials 11–13; infralabials 9–10; dorsal tubercles at midbody in 4–6 irregular rows; ventral scales 52–68; subdigital lamellae on the fourth toe 17–19; precloacal and femoral pores absent in both sexces; enlarged precloacal scales 22–27 in males and absent in females; 7–11 enlarged femoral scales from each thigh; subcaudals transversely enlarged; small, irregularly shaped dark brown blotches on top of head; nuchal band thin and interrupted; dorsal pattern with four irregularly shaped bands; short stripes on the neck. The morphological distinctiveness of the new species is supported by its non-overlapping position in multivariate space based on multiple factor analysis and principal component analysis.
References
- Barraclough, T.G., Birky, C.W. Jr. & Burt, A. (2003) Diversification in sexual and asexual organisms. International Journal of Organic Evolution, 57, 2166–2172. https://doi.org/10.1554/02-339
- Bauer, A.M., Pauwels, O.S.G. & Chanhome, L. (2002) A New Species of Cave–dwelling Cyrtodactylus (Squamata: Gekkonidae) from Thailand. Natural History Journal of Chulalongkorn University, 2 (2), 19–29. https://doi.org/10.58837/tnh.2.2.102853
- Chan, K.O. & Grismer, L.L. (2022) GroupStruct: An R package for allometric size correction. Zootaxa, 5124 (4), 471–482. https://doi.org/10.11646/zootaxa.5124.4.4
- David, P., Teynié, A. & Ohler, A. (2004) A new species of Cyrtodactylus Gray, 1827 (Reptilia: Squamata: Gekkonidae) from southern Laos. Raffles Bulletin of Zoology, 52, 621–627.
- de Queiroz, K. (2007) Species concepts and species delimitation. Systematic Biology, 56 (6), 879–886. https://doi.org/10.1080/10635150701701083
- Do, Q.H., Phung, T.M., Ngo, H.T., Le, M.D., Ziegler, T., Pham, C.T. & Nguyen, T. (2021) A new species of Cyrtodactylus (Squamata: Gekkonidae) from Ninh Thuan Province, southern Vietnam. Zootaxa, 4999 (1), 58–76. https://doi.org/10.11646/zootaxa.4999.1.4
- Drummond, A.J., Suchard, M.A., Xie, D. & Rambaut, A. (2012) Bayesian Phylogenetics with BEAUti and BEAST 1.7. Molecular Biology and Evolution, 29 (8), 1969–1973. https://doi.org/10.1093/molbev/mss075
- Frost, D.R. & Hillis, D.M. (1990) Species in concept and practice: Herpetological application. Herpetologica, 46, 87–104.
- Frost, D.R. & Kluge, A.G. (1994) A consideration of the epistemology in systematic biology, with special reference to species. Cladistics, 10 (3), 259–294. https://doi.org/10.1111/j.1096-0031.1994.tb00178.x
- Gray, J.E. (1827) A Synopsis of the genera of saurian reptiles in which some new genera are indicated and the others reviewed by actual Examination. Philosophical Magazine London, 2, 54–58. https://doi.org/10.1080/14786442708675620
- Geissler, P., Nazarov, R.A., Orlov, N.L., Phung, T.M., Nguyen, T.Q. & Ziegler, T. (2009) A new species of the Cyrtodactylus irregularis complex (Squamata, Gekkonidae) from southern Vietnam. Zootaxa, 2161 (1), 20–32. https://doi.org/10.11646/zootaxa.2161.1.2
- Grismer, L.L., Wood, P.L. & Ngo, V.T. & Murdoch, M.L. (2015) The systematics and independent evolution of cave eco-morphology in distantly related clades of bent-toed geckos (genus Cyrtodactylus Gray, 1827) from the Mekong Delta and islands in the Gulf of Thailand. Zootaxa, 3980 (1), 106–126. https://doi.org/10.11646/zootaxa.3980.1.6
- Grismer, L.L., Wood, P.L. Jr., Le, M.D., Quah, E.S.H. & Grismer, J.L. (2020) Evolution of habitat preference in 243 species of bent-toed geckos (genus Cyrtodactylus Gray, 1827) with a discussion of karst habitat conservation. Ecology and Evolution, 10 (24), 13717–13730. https://doi.org/10.1002/ece3.6961
- Grismer, L.L., Wood, P.L. Jr., Poyarkov, N.A., Le, M.D., Kraus, F., Agarwal, I., Oliver, P.M., Nguyen, S.N., Nguyen, T.Q., Karunarathna, S., Welton, L.J., Stuart, B.L., Luu, V.Q., Bauer, A.M., O’Connell, K.A., Quah, E.S.H., Chan, K.O., Ziegler, T., Ngo, H., Nazarov, R.A., Aowphol, A., Chomdej, S., Suwannapoom, C., Siler, C.D., Anuar, S. & Grismer, J.L. (2021) Phylogenetic partitioning of the third-largest vertebrate genus in the world, Cyrtodactylus Gray, 1827 (Reptilia; Squamata; Gekkonidae) and its relevance to taxonomy and conservation. Vertebrate Zoology, 71, 101–154. https://doi.org/10.3897/vz.71.e59307
- Grismer, L.L., Sinovas, P., Quah, E.S.H., Thi, S., Chourn, P., Chhin, S., Hun, S., Cobos, A., Geissler, P., Ching, C. & Murdoch, M.L. (2025) A new species of lowland karst-dwelling Slender Gecko Hemiphyllodactylus Bleeker, 1860 (Squamata: Gekkonidae) from a karstic archipelago in western Cambodia. Zootaxa, 5569 (2), 253–281. https://doi.org/10.11646/zootaxa.5569.2.3
- Grismer, L.L., Wood, P.L. Jr., Thura, M.K., Zin, T., Quah, E.S.H., Murdoch, M.L., Grismer, M.S., Lin, A., Kyaw, H. & Lwin, N. (2018) Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in eastcentral and southern Myanmar demonstrate high localized diversity and unprecedented microendemism. Zoological Journal of the Linnean Society, 182, 862–959. https://doi.org/10.1093/zoolinnean/zlx057
- Heidrich, A., Rösler, H., Vu, N.T., Böhme, W. & Ziegler, T. (2007) Another new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha-Ke Bang National Park, central Truong Son, Vietnam. Zootaxa, 1445 (1), 35–48. https://doi.org/10.11646/zootaxa.1445.1.3
- Hillis, D.M. (2019) Species delimitation in herpetology. Journal of Herpetology, 53 (1), 3–12. https://doi.org/10.1670/18-123
- Hoang, D.T., Chernomor, O., von Haeseler, A., Minh, B.Q. & Vinh, L.S. (2018) UFBoot2: Improving the ultrafast bootstrap approximation. Molecular Biology and Evolution, 35 (2), 518–522. https://doi.org/10.1093/molbev/msx281
- Huelsenbeck, J.P., Ronquist, F., Nielsen, R. & Bollback, J.P. (2001) Bayesian inference of phylogeny and its impact on evolutionary biology. Science, 294 (5550), 2310–2314. https://doi.org/10.1126/science.1065889
- Husson, F., Josse, J., Le, S. & Mazet, J. (2017) FactoMineR: Exploratory Data Analysis and Data Mining. R package version 1.36.
- Jombart, T., Devillard, S. & Balloux, F. (2010) Discriminant analysis of principal components: A new method for the analysis of genetically structured populations. BMC Genetics, 11 (1), 1–15. https://doi.org/10.1186/1471-2156-11-94
- Kalyaanamoorthy, S., Minh, B.Q., Wong, T.K., von Haeseler, A. & Jermiin, L.S. (2017) ModelFinder: Fast model selection for accurate phylogenetic estimates. Nature Methods, 14 (6), 587–589. https://doi.org/10.1038/nmeth.4285
- Kassambara, A. & Mundt, F. (2017) factoextra: Extract and Visualize the Result of Multivariate Data Analyses. R package Version 1.0.5.999. [program]
- Kumar, S., Stecher, G. & Tamura, K. (2016) MEGA7: Molecular evolutionary genetics analysis version 7.0 for bigger datasets. Molecular Biology and Evolution, 33 (7), 1870–1874. https://doi.org/10.1093/molbev/msw054
- Lleonart, J., Salat, J. & Torres, G.J. (2000) Removing allometric effects of body size in morphological analysis. Journal of Theoretical Biology, 205 (1), 85–93. https://doi.org/10.1006/jtbi.2000.2043
- Luu, V.Q., Calame, T., Nguyen, T.Q., Bonkowski, M. & Ziegler, T. (2015) A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa, 4058 (3), 388–402. https://doi.org/10.11646/zootaxa.4058.3.6
- Luu, V.Q., Bonkoski, M., Nguyen, T.Q., Le, M.D., Schneider, N., Ngo, H.T. & Ziegler, T. (2016) Evolution in karst massifs: cryptic diversity among bent-toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos. Zootaxa, 4107 (2), 101–140. https://doi.org/10.11646/zootaxa.4107.2.1
- Luu, V.Q., Dung, T.V., Nguyen, T.Q., Le, M.D. & Ziegler, T. (2017) A new species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from Gia Lai Province, Central Highlands of Vietnam. Zootaxa, 4362 (3), 385–404. https://doi.org/10.11646/zootaxa.4362.3.4
- Luu, V.Q., Van, O.L., Hoang, T.T., Van, T.P., Le Duc, O., Bordes, C., Leprince, B., Amori, G. & Luiselli, L. (2020) Ecological characteristics of a recently described, critically endangered gecko species, endemic to Central Highland, Vietnam. Tropical Zoology, 33 (2). [published online] https://doi.org/10.4081/tz.2020.71
- Luu, V.Q. (2018) Cyrtodactylus gialaiensis. The IUCN Red List of Threatened Species, 2018, e.T125554212A125554215. Available from: https://doi.org/10.2305/IUCN.UK.2018-2.RLTS.T125554212A125554215.en/ (accessed 8 April 2025)
- Luu, V.Q., Nguyen, T.H., Le, M.D., Grismer, J.L., Ha, H.B., Sitthivong, S., Hoang, T.T. & Grismer, L.L. (2023) Two new species of Dixonius from Vietnam and Laos with a discussion of the taxonomy of Dixonius (Squamata, Gekkonidae). ZooKeys, 1163, 143–176. https://doi.org/10.3897/zookeys.1163.101230
- Macey, J.R., Larson, A., Ananjeva, N.B., Fang, Z. & Papenfuss, T.J. (1997) Two novel gene orders and the role of light-strand replication in rearrangement of the vertebrate mitochondrial genome. Molecular Biology and Evolution, 14 (1), 91–104. https://doi.org/10.1093/oxfordjournals.molbev.a025706
- McCoy, M.W., Bolker, B.M., Osenberg, C.W., Miner, B.G. & Vonesh, J.R. (2006) Size correction: Comparing morphological traits among populations and environments. Oecologia, 148 (4), 547–554. https://doi.org/10.1007/s00442-006-0403-6
- Miller, M.A., Pfeiffer, W. & Schwartz, T. (2010) Creating the CIPRES Science Gateway for inference of large phylogenetic trees. Proceedings of the Gateway Computing Environments Workshop (GCE), New Orleans, Louisiana, 14 November 2010, 1–8. https://doi.org/10.1109/GCE.2010.5676129
- Minh, Q., Nguyen, M.A.T. & von Haeseler, A. (2013) Ultrafast approximation for phylogenetic bootstrap. Molecular Biology and Evolution, 30 (5), 1188–1195. https://doi.org/10.1093/molbev/mst024
- Nazarov, R.A., Orlov, N.L., Nguyen, N.S. & Ho, T.C. (2008) Taxonomy of naked-toe geckos Cyrtodactylus irregularis complex of South Vietnam and description of a new species from Chu Yang Sin Natural Park (Krong Bong District, Dac Lac Province), Vietnam. Russian Journal of Herpetology, 15, 141–156.
- Nazarov, R., Poyarkov, N.A., Orlov, N.L., Phung, T.M., Nguyen, T.T., Hoang, D.M. & Ziegler, T. (2012) Two new cryptic species of the Cyrtodactylus irregularis complex (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 3302 (1), 1–24. https://doi.org/10.11646/zootaxa.3302.1.1
- Neang, T., Henson, A. & Stuart, B.L. (2020) A new species of Cyrtodactylus (Squamata, Gekkonidae) from Cambodia’s Prey Lang Wildlife Sanctuary. ZooKeys, 926, 133–158. https://doi.org/10.3897/zookeys.926.48671
- Ngo, H.T., Do, Q.H., Pham, C.T., Luu, V.Q., Grismer, L.L., Ziegler, T., Nguyen, V.T.H., Nguyen, T.Q. & Le, M.D. (2022) How many more species are out there? Current taxonomy seriously underestimates the diversity of bent-toed geckos (genus Cyrtodactylus) in Laos and Vietnam. ZooKeys, 1097, 135–152. https://doi.org/10.3897/zookeys.1097.78127
- Ngo, H.T., Hormann, H., Le, M.D., Pham, C.T., Phung, T.M., Do, D.T., Ostrowski, S., Nguyen, T.Q. & Ziegler, T. (2023) The discovery of two new species in the Cyrtodactylus irregularis group highlights that hidden diversity remains in the largest clade of the mega-diverse genus Cyrtodactylus. European Journal of Taxonomy, 875, 70–100. https://doi.org/10.5852/ejt.2023.875.2141
- Ngo, T.V. (2008) Two new cave-dwelling species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Southwestern Vietnam. Zootaxa, 1909 (1), 37–51. https://doi.org/10.11646/zootaxa.1909.1.4
- Ngo, T.V. (2013) Cyrtodactylus dati, a new forest dwelling Bent-toed Gecko (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 3616 (2), 151–164. https://doi.org/10.11646/zootaxa.3616.2.4
- Ngo, T.V. & Bauer, A.M. (2008) Descriptions of two new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) endemic to southern Vietnam. Zootaxa, 1715 (1), 27–42. https://doi.org/10.11646/zootaxa.1715.1.2
- Ngo, T.V. & Chan, K.O. (2010) A new species of Cyrtodactylus Gray, 1826 (Squamata: Gekkonidae) from Khanh Hoa province, southern Vietnam. Zootaxa, 2504 (1), 47–60. https://doi.org/10.11646/zootaxa.2504.1.4
- Ngo, T.V. & Chan, O.K. (2011) A new krastic cave-dwelling Cyrtodactylus Gray (Squamata: Gekkonidae) from Northern Vietnam. Zootaxa, 3125 (1), 51–63. https://doi.org/10.11646/zootaxa.1909.1.4
- Ngo, T.V. & Grismer, L.L. (2012) A new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from Tho Chu Island, southwestern Vietnam. Zootaxa, 3228 (1), 48–60. https://doi.org/10.11646/zootaxa.3228.1.2
- Ngo, T.V., Grismer, L.L. & Grismer, J.L. (2010) A new species of Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) in Phu Quoc National Park, Kien Giang Biosphere Reserve, Southwestern Vietnam. Zootaxa, 2604 (1), 37–51. https://doi.org/10.11646/zootaxa.2604.1.3
- Ngo, T.V., Grismer, L.L., Pham, T.H. & Wood, P.L. Jr. (2020) A new endemic insular Bent-toed Gecko (Squamata: Gekkonidae: Cyrtodactylus) from Quang Nam Province, Central Vietnam. Zootaxa, 4766 (2), 389–400. https://doi.org/10.11646/zootaxa.4766.2.7
- Nguyen, A.T., Duong, T.V., Grismer, L.L. & Poyarkov, N.A. (2021) A new granite cave-dwelling Bent-toed Gecko from Vietnam of the Cyrtodactylus irregularis group (Squamata: Gekkonidae) and a discussion on cave ecomorphology. Vertebrate Zoology, 71, 155–174. https://doi.org/10.3897/vz.71.e60225
- Nguyen, L.T., Schmidt, H.A., von Haeseler, A. & Minh, B.Q. (2015) IQ-TREE: A fast and effective stochastic algorithm for estimating maximum likelihood phylogenies. Molecular Biology and Evolution, 32 (1), 268–274. https://doi.org/10.1093/molbev/msu300
- Nguyen, T.Q. (2023) Cyrtodactylus gialaiensis. The Vietnam Red List of Threatened Species, RT22. Available from: http://vnredlist.vast.vn/cyrtodactylus-gialaiensis/ (accessed 8 April 2025)
- Nguyen, S.N., Orlov, N.L. & Darevsky, I.S. (2006) Descriptions of two new species of the genus Cyrtodactylus Gray, 1827 (Squamata: Sauria: Gekkonidae) from southern Vietnam. Russian Journal of Herpetology, 13 (3), 215–226. https://doi.org/10.30906/1026-2296-2006-13-3-215-226
- Nguyen, S.N., Le, T.T., Tran, T.A.D., Orlov, N.L., Lathrop, A., MacCulloch, R.D., Le, T.D.T., Jun, J-Q., Nguyen, L.T., Nguyen, T.T., Hoang, D.D., Che, J., Murphy, R.Q. & Zhang, Y.P. (2013) Phylogeny of the Cyrtodactylus irregularis species complex (Squamata: Gekkonidae) from Vietnam with the description of two new species. Zootaxa, 3737 (4), 399–414. https://doi.org/10.11646/zootaxa.3737.4.4
- Nguyen, S.N., Nguyen, L.T., Nguyen, V.D.H. & Murphy, R.W. (2017b) Genetic diversity, phylogeny, and altitudinal distribution of geckos genus Cyrtodactylus on Hon Ba Mountain, Khanh Hoa Province, with note on the subcaudals of C. yangbayensis. Journal of Biotechnology, 15 (3A), 55–62.
- Orlov, N.L., Nguyen, T.Q., Nazarov, A.R., Ananjeva, B.N. & Nguyen, S.N. (2007) A new species of the genus Cyrtodactylus Gray, 1827 and redescription of Cyrtodactylus paradoxus (Darevsky et Szczerbak, 1997) [Squamata: Sauria: Gekkonidae] from South Vietnam. Russian Journal of Herpetology, 14, 145–152.
- Ostrowski, S., Le, M.D., Ngo, H.T., Pham, C.T., Nguyen, T.Q. & Ziegler, T. (2020) A new species of Cyrtodactylus (Squamata: Gekkonidae) from southern Vietnam. Zootaxa, 4789 (1), 171–203. https://doi.org/10.11646/zootaxa.4789.1.5
- Ostrowski, S., Le, M.D., Ngo, H.T., Pham, C.T., Phung, T.M., Nguyen, T.Q. & Ziegler, T. (2021) A new Cyrtodactylus (Squamata: Gekkonidae) from Binh Thuan Province, southern Vietnam. European Journal of Taxonomy, 731 (1), 47–70. https://doi.org/10.5852/ejt.2021.731.1023
- Pagès, J. (2015) Multiple Factor Analysis by Example Using R. CRC Press, New York, New York, 272 pp. https://doi.org/10.1201/b17700
- Pauwels, O.S.G., Nazarov, R.A., Bobrov, V.V. & Poyarkov, N.A. (2018) Taxonomic status of two populations of bent-toed geckos of the Cytrodactylus irregularis complex (Squamata: Gekkonidae) with description of a new species from Nui Chua National Park, southern Vietnam. Zootaxa, 4403 (2), 307–335. https://doi.org/10.11646/zootaxa.4403.2.5
- Phung, T.M., van Schingen, M., Ziegler, T. & Nguyen, T.Q. (2014) A third new Cyrtodactylus (Squamata: Gekkonidae) from Ba Den Mountain, Tay Ninh Province, southern Vietnam. Zootaxa, 3764 (3), 347–363. https://doi.org/10.11646/zootaxa.3764.3.5
- R Core Team (2021) R: A language and environment for statistical computing. R foundation for statistical computing, Vienna. Available from: https://www.R-project.org (accessed 15 July 2025)
- Rambaut, A. & Drummond, A.J. (2013) TreeAnnotator. Version 1.7.3 MCMC Output Analysis. Available from: https://beast.community/treeannotator (accessed 15 January 2023)
- Rambaut, A., Suchard, M.A., Xie, D. & Drummond, A.J. (2014) Tracer. Version 1.7. Available from: http://tree.bio.ed.ac.uk/software/tracer/ (accessed 15 January 2023)
- Reist, J.D. (1986) An empirical evaluation of coefficients used in residual and allometric adjustment of size covariation. Canadian Journal of Zoology, 64 (6), 1363–1368. https://doi.org/10.1139/z86-203
- Ronquist, F., Teslenko, M., van der Mark, P., Ayres, D.L., Darling, A., Höhna, S., Larget, B., Liu, L., Suchard, M.A. & Huelsenbeck, J.P. (2012) MrBayes 3.2: efficient Bayesian phylogenetic inference and model choice across a large model space. Systematic Biology, 61 (3), 539–542. https://doi.org/10.1093/sysbio/sys029
- Rösler, H., Vu, T.N., Nguyen, T.Q., Ngo, V.T. & Ziegler, T. (2008) A new Cyrtodactylus (Squamata: Gekkonidae) from central Vietnam. Hamadryad, 32, 125–140.
- Schneider, N., Phung, T.M., Le, M.D., Nguyen, T.Q. & Ziegler, T. (2014) A new Cyrtodactylus (Squamata: Gekkonidae) from Khanh Hoa Province, southern Vietnam. Zootaxa, 3785 (4), 518–532. https://doi.org/10.11646/zootaxa.3785.4.2
- Smith, M.A. (1921) New or little-known Reptiles and Batrachians from Southern Annam (Indochina). Proceedings of the Zoological Society of London, 1921, 423–440. https://doi.org/10.1111/j.1096-3642.1921.tb03271.x
- Thorpe, R.S. (1975) Quantitative handling of characters useful in snake systematics with particular reference to interspecific variation in the Ringed Snake Natrix natrix (L.). Biological Journal of the Linnean Society, 7 (1), 27–43. https://doi.org/10.1111/j.1095-8312.1975.tb00732.x
- Thorpe, R.S. (1983) Phylogenetic analysis of range expansion in the grass snake: reticulate evolution: primary and secondary contact zones. Numerical Taxonomy: Proceedings of a NATO Advanced Study Institute. In: Felsenstein, J. (Ed.), NATO Advanced Study Institute. Series G (Ecological Sciences). No. 1. Springer‑Verlag, Berlin, Heidelberg and New York pp. 464–468. https://doi.org/10.1007/978-3-642-69024-2_47
- Trifinopoulos, J., Nguyen, L.T., von Haeseler, A. & Minh, B.Q. (2016) W-IQ-TREE: A fast online phylogenetic tool for maximum likelihood analysis. Nucleic Acids Research, 44 (W1), W232–W235. https://doi.org/10.1093/nar/gkw256
- Turan, C. (1999) A note on the examination of morphometric differentiation among fish populations: The Truss System. Turkish Journal of Zoology, 23, 259–263.
- Uetz, P., Freed, P. & Hosek, J. (2025) The Reptile Database. Available from: http://www.reptile-database.org/ (accessed 8 April 2025)
- Wilcox, T.P., Zwickl, D.J., Heath, T.A. & Hillis, D.M. (2002) Phylogenetic relationships of the Dwarf Boas and a comparison of Bayesian and bootstrap measures of phylogenetic support. Molecular Phylogenetics and Evolution, 25 (2), 361–371. https://doi.org/10.1016/S1055-7903(02)00244-0
- Ziegler, T., Rösler, H., Herrmann, H.-W. & Vu, T.N. (2002) Cyrtodactylus phongnhakebangensis sp. n., ein neuer Bogenfingergecko aus dem annamitischen Karstwaldmassiv, Vietnam. Herpetofauna, 24 (141), 11–25.
- Ziegler, T., Nazarov, R., Orlov, N., Nguyen, T.Q., Vu, T.N., Dang, K.N., Dinh, T.H. & Schmitz, A. (2010) A third new Cyrtodactylus (Squamata: Gekkonidae) from Phong Nha-Ke Bang National Park, Truong Son Range, Vietnam. Zootaxa, 2413 (1), 20–36. https://doi.org/10.11646/zootaxa.2413.1.2
- Ziegler, T., Phung, T.M., Le, M.D. & Nguyen, T.Q. (2013) A new Cyrtodactylus (Squamata: Gekkonidae) from Phu Yen Province, southern Vietnam. Zootaxa, 3686 (4), 432–446. https://doi.org/10.11646/zootaxa.3686.4.2