Skip to main content Skip to main navigation menu Skip to site footer
Type: Review
Published: 2025-04-29
Page range: 195–234
Abstract views: 58
PDF downloaded: 13

Cretaceous Spanish amber: History, research and checklist of taxa

Instituto Geológico y Minero de España (IGME), CSIC, C/ Cirilo Amorós 42, 46004 Valencia, Spain
Departamento de Biodiversidad, Ecología y Evolución, Facultad de Biología, Universidad Complutense, 28040 Madrid, Spain
Museo de Ciencias Naturales de Álava, C/ Siervas de Jesús, 24. 01001 Vitoria-Gasteiz (Álava), Spain
Departament de Dinàmica de la Terra i de l’Oceà, Facultat de Ciències de la Terra, Universitat de Barcelona, 08028 Barcelona, Spain; Institut de Recerca de la Biodiversitat (IRBio), Universitat de Barcelona, Barcelona, Spain
bioinclusions described taxa holotypes type localities Mesozoic

Abstract

The present work is a checklist of taxa described or cited from bioinclusions preserved in Spanish amber outcrops (nine type localities). All these bioinclusions are Albian in age (ca. 110–102 Ma), except for a few from a Cenomanian outcrop. All the listed taxa are arthropods (mostly insects), with the exception of some plant parataxa and plant palynomorphs. The present checklist of taxa has been elaborated 30 years after the first bioinclusion was discovered in Spain, thus virtually it spans over 30 years of study by a Spanish team (AMBERIA) thanks to a series of national projects funded by the Spanish Ministry of Science and other public institutions as the Natural Science Museum of Álava (Provincial Council of Álava), and the Government of Cantabria together with the semi-public enterprise EL SOPLAO S.L. The list comprises 7 classes (or similar rank), 24 orders (or similar rank), 101 families, 169 genera and 186 species (including the plant parataxa). Eight families, 104 genera (implicating a total of 104 type species) and 184 species were described based on 184 holotypes from Spanish amber. We provide all the relevant data for each taxon, focusing on new families, genera and species described based on Spanish bioinclusions, pretending to be as useful and readable as possible.

References

  1. Ábalos, B. (2016) Geologic map of the Basque‑Cantabrian Basin and a new tectonic interpretation of the Basque Arc. International Journal of Earth Sciences, 105 (8), 2327–2354. https://doi.org/10.1007/s00531-016-1291-6
  2. Alonso, J., Arillo, A., Barrón, E., Corral, J.C., Grimalt, J., López, J.F., López, R., Martínez-Delclòs, X., Ortuño, V., Peñalver, E. & Trincão, P.R. (2000) A new fossil resin with biological inclusions in lower Cretaceous deposits from Álava (Northern Spain, Basque-Cantabrian Basin). Journal of Paleontology, 74, 158–178. https://doi.org/10.1666/0022-3360(2000)074<0158:ANFRWB>2.0.CO;2
  3. Álvarez-Parra, S. (2023) Yacimientos de ámbar del Cretácico de la Cuenca del Maestrazgo: tafonomía, bioinclusiones y paleoecología. Ph.D. thesis. University of Barcelona (unpublished), 522 pp. https://diposit.ub.edu/dspace/handle/2445/193847
  4. Álvarez-Parra, S., Delclòs, X., Solórzano-Kraemer, M.M., Alcalá, L. & Peñalver, E. (2020) Cretaceous amniote integuments recorded through a taphonomic process unique to resins. Scientific Reports, 10, 19840. https://doi.org/10.1038/s41598-020-76830-8
  5. Álvarez-Parra, S., Engel, M.S., Peñalver, E. & Azar, D. (2024a) The aculeate wasp family Sierolomorphidae (Hymenoptera) in the Early Cretaceous. Insect Systematics and Diversity, 8 (4), 1–7. https://doi.org/10.1093/isd/ixae020
  6. Álvarez-Parra, S., Jouault, C. & Azar, D. (2024b) A braconid wasp (Hymenoptera: Braconidae) with second recurrent vein in forewing from Lower Cretaceous Spanish amber. Palaeoentomology, 7 (4), 479–490. https://doi.org/10.11646/palaeoentomology.7.4.7
  7. Álvarez-Parra, S., Peñalver, E., Delclòs, X. & Engel, M.S. (2022a) A braconid wasp (Hymenoptera, Braconidae) from the Lower Cretaceous amber of San Just, eastern Iberian Peninsula. ZooKeys, 1103, 65–78. https://doi.org/10.3897/zookeys.1103.83650
  8. Álvarez-Parra, S., Peñalver, E., Nel, A. & Delclòs, X. (2022b) New barklice (Psocodea: Trogiomorpha) from Lower Cretaceous Spanish amber. Papers in Palaeontology, 8 (3), e1436. https://doi.org/10.1002/spp2.1436
  9. Álvarez-Parra, S., Peñalver, E., Nel, A. & Delclòs, X. (2023) Barklice (Insecta: Psocodea) from Early Cretaceous resiniferous forests of Iberia (Spanish amber): New Troctomorpha and a possible Psocomorpha. Cretaceous Research, 148, 105544. https://doi.org/10.1016/j.cretres.2023.105544
  10. Álvarez-Parra, S., Peñalver, E., Nel, P., Nel, A. & Delclòs, X. (2021a) A representative of the controversial order †Lophioneurida (Insecta: Thripida) in Albian Spanish amber. In: Vlachos, E., Penélope Cruzado, C., Crespo, V.D., Ríos Ibañez, M., Arnal, F.A.M., Herraiz, J.L., Gascó-Lluna, F., Guerrero-Arenas, R. & Ferrón, H.G. (Eds.). Book of Abstracts of the 3rd Palaeontological Virtual Congress, 187.
  11. Álvarez-Parra, S., Pérez-de la Fuente, R., Peñalver, E., Barrón, E., Alcalá, L., Pérez-Cano, J., Martín-Closas, C., Trabelsi, K., Meléndez, N., López Del Valle, R., Lozano, R.P., Peris, D., Rodrigo, A., Sarto i Monteys, V., Bueno-Cebollada, C.A., Menor-Salván, C., Philippe, M., Sánchez-García, A., Peña-Kairath, C., Arillo, A., Espílez, E., Mampel, L. & Delclòs, X. (2021b) Dinosaur bonebed amber from an original swamp forest soil. eLife, 10, e72477. https://doi.org/10.7554/eLife.72477
  12. Antolín-Tomás, B., Liesa, C.L., Casas, A. & Gil-Peña, I. (2007) Geometry of fracturing linked to extension and basin formation in the Maestrazgo Basin (Eastern Iberian Chain, Spain). Revista de la Sociedad Geológica de España, 20 (3-4), 351–365.
  13. Arbizu, M., Bernárdez, E., Peñalver, E. & Prieto, M.A. (1999) El ámbar de Asturias. Estudios del Museo de Ciencias Naturales de Álava, 14 (nº especial 2), 245–254.
  14. Arillo, A. (1996) Los insectos en ámbar. Boletín de la Sociedad Aragonesa de Entomología, 16, 147–149.
  15. Arillo, A. & Mostovski, M.B. (1999) A new genus of Prioriphorinae (Diptera, Phoridae) from the Lower Cretaceous amber of Alava (Spain). Studia dipterologica, 6 (2), 251–255.
  16. Arillo, A. & Nel, A. (2000) Two new fossil cecidomyiids flies from the Lower Cretaceous amber of Alava (Spain) (Diptera, Cecidomyiidae). Bulletin de la Société entomologique de France, 105 (3), 285–288. https://doi.org/10.3406/bsef.2000.16673
  17. Arillo, A. & Subías, L.S. (2000) A new fossil oribatid mite, Archaeorchestes minguezae gen. nov. sp. nov. from the Spanish Lower Cretaceous amber. Description of a new family, Archaeorchestidae (Acariformes, Oribatida, Zetorchestoidea). Mitteilungen aus dem Geologisch-Paläontologischen Institut der Universität Hamburg, 84, 231–236.
  18. Arillo, A. & Subías, L.S. (2002) Second fossil oribatid mite from the Spanish Lower Cretaceous amber. Eupterotegaeus bitranslamellatus n. sp. (Acari, Oribatida, Cepheidae). Acarologia, 42 (4), 403–406.
  19. Arillo, A., Blagoderov, V. & Peñalver, E. (2018) Early Cretaceous parasitism in amber: A new species of Burmazelmira fly (Diptera: Archizelmiridae) parasitized by a Leptus sp. mite (Acari, Erythraeidae). Cretaceous Research, 86, 24–32. https://doi.org/10.1016/j.cretres.2018.02.006
  20. Arillo, A., Peñalver, E. & Delclòs, X. (2008a) Microphorites (Diptera: Dolichopodidae) from the Lower Cretaceous amber of San Just (Spain), and the co-occurrence of two ceratopogonid species in Spanish amber deposits. Zootaxa, 1920 (1), 29–40. https://doi.org/10.11646/zootaxa.1920.1.2
  21. Arillo, A., Peñalver, E. & García-Gimeno, V. (2009a) First fossil Litoleptis (Diptera: Spaniidae) from the Lower Cretaceous amber of San Just (Teruel Province, Spain). Zootaxa, 2026 (1), 33–39. https://doi.org/10.11646/zootaxa.2026.1.3
  22. Arillo, A., Peñalver, E., Pérez-de la Fuente, R., Delclòs, X., Criscione, J., Barden, P.M., Riccio, M.L. & Grimaldi, D.A. (2015) Long-proboscid brachyceran flies in Cretaceous amber (Diptera: Stratiomyomorpha: Zhangsolvidae). Systematic Entomology, 40, 242–267. https://doi.org/10.1111/syen.12106
  23. Arillo, A., Subías, L.S. & Álvarez-Parra, S. (2022) First fossil record of the oribatid family Liacaridae (Acariformes: Gustavioidea) from the lower Albian amber-bearing site of Ariño (eastern Spain). Cretaceous Research, 131, 105087. https://doi.org/10.1016/j.cretres.2021.105087
  24. Arillo, A., Subías, L.S. & Peñalver, E. (2020) A new species of fossil Oribatid mite (Acariformes, Oribatida: Caleremaeidae) from a new Cretaceous amber outcrop in Asturias, Spain. Cretaceous Research, 109, 104382. https://doi.org/10.1016/j.cretres.2020.104382
  25. Arillo, A., Subías, L.S. & Sánchez-García, A. (2016) New species of fossil oribatid mites (Acariformes, Oribatida), from the Lower Cretaceous amber of Spain. Cretaceous Research, 63, 68–76. https://doi.org/10.1016/j.cretres.2016.02.009
  26. Arillo, A., Subías, L.S. & Shtanchaeva, U. (2008b) A new fossil oribatid mite, Ommatocepheus nortoni sp. nov. (Acariformes, Oribatida, Cepheidae) from a new outcrop of Lower Cretaceous Álava amber (northern Spain). Systematic & Applied Acarology, 13 (3-4), 252–255. https://doi.org/10.11158/saa.13.3.13
  27. Arillo, A., Subías, L.S. & Shtanchaeva, U. (2009b) A new fossil species of oribatid mite, Ametroproctus valeriae sp. nov. (Acariformes, Oribatida, Ametroproctidae), from the Lower Cretaceous amber of San Just, Teruel Province, Spain. Cretaceous Research, 30, 322–324. https://doi.org/10.1016/j.cretres.2008.07.013
  28. Arillo, A., Subías, L.S. & Shtanchaeva, U. (2010) A new genus and species of oribatid mite, Cretaceobodes martinezae gen. et sp. nov., from the Lower Cretaceous amber of San Just (Teruel Province, Spain) (Acariformes, Oribatida, Otocepheidae). Paleontological Journal, 44, 287–290. https://doi.org/10.1134/S003103011003007X
  29. Arillo, A., Subías, L.S. & Shtanchaeva, U. (2012) A new species of fossil oribatid mite (Acariformes, Oribatida, Trhypochthoniidae) from the Lower Cretaceous amber of San Just (Teruel Province, Spain). Systematic & Applied Acarology, 17 (1), 106–112. https://doi.org/10.11158/saa.17.1.16
  30. Bahamonde, J. (1984) Análisis sedimentológico de las facies detríticas del Cretácico de los alrededores de Oviedo. Tesis de Licenciatura. Universidad de Oviedo. (unpublished), 60 pp.
  31. Barnolas, A. & Pujalte, V. (2004) La Cordillera Pirenaica: definición, límites y división. In: Vera, J.A. (Ed.), Geología de España. Sociedad Geológica de España. Instituto Geológico y Minero de España, Madrid, 233–241.
  32. Barrón, E., Peyrot, D., Bueno-Cebollada, C.A., Kvaček, J., Álvarez-Parra, S., Altolaguirre, Y. & Meléndez, N. (2023) Biodiversity of ecosystems in an arid setting: The late Albian plant communities and associated biota from eastern Iberia. PLoS ONE, 18 (3), e0282178. hhttps://doi.org/10.1371/journal.pone.0282178
  33. Barrón, E., Peyrot, D., Rodríguez-López, J.P., Meléndez, N., López del Valle, R., Najarro, M., Rosales, I. & Comas-Rengifo, Mª J. (2015) Palynology of Aptian and upper Albian (Lower Cretaceous) amber-bearing outcrops of the southern margin of the Basque-Cantabrian basin (northern Spain). Cretaceous Research, 52, 292–312. https://doi.org/10.1016/j.cretres.2014.10.003
  34. Baz, A. & Ortuño, V.M. (2000) Archaeatropidae, a new family of Psocoptera from the Cretaceous amber of Alava, northern Spain. Annals of the Entomological Society of America, 93 (3), 367–373. https://doi.org/10.1603/0013-8746(2000)093[0367:AANFOP]2.0.CO;2
  35. Baz, A. & Ortuño, V.M. (2001a) New genera and species of empheriids (Psocoptera: Empheriidae) from the Cretaceous amber of Álava, northern Spain. Cretaceous Research, 22 (5), 575–584. https://doi.org/10.1006/cres.2001.0275
  36. Baz, A. & Ortuño, V.M. (2001b) A new electrentomoid psocid (Psocoptera) from the Cretaceous amber of Álava (Northern Spain). Zoosystematics and Evolution, 48 (1), 27–32. https://doi.org/10.1002/mmnd.4800480104
  37. Bernárdez, E. (1994) Unidades litoestratigráficas del Cretácico de la Depresión Central Asturiana. Cuadernos de Geología Ibérica, 18, 11–25.
  38. Bernárdez, E. (2005) Discusión sobre la Revisión y síntesis litoestratigráfica del sector occidental de la cuenca cretácica de Asturias. Réplica. Trabajos de Geología, Universidad de Oviedo, 25, 117–137.
  39. Blagoderov, V. & Arillo, A. (2002) New Sciaroidea (Insecta: Diptera) in Lower Cretaceous amber from Spain. Studia dipterologica, 9 (1), 31–40.
  40. Borkent, A. (2019) The phylogenetic relationships of Cretaceous biting midges, with a key to all known genera (Diptera: Ceratopogonidae). American Museum Novitates, 3921, 1–48. https:/doi.org/10.1206/3921.1
  41. Bover-Arnal, T., Moreno-Bedmar, J.A., Frijia, G., Pascual-Cebrian, E. & Salas, R. (2016) Chronostratigraphy of the Barremian-Early Albian of the Maestrat Basin (E Iberian Peninsula): integrating strontium-isotope stratigraphy and ammonoid biostratigraphy. Newsletters on Stratigraphy, 49, 41–68. https://doi.org/10.1127/nos/2016/0072
  42. Casal, G. (1762) Succini Asturici, à Doctore Gafsar Casal, Almae Eclesiae Cathedralis Ovetensis Medico, reperti, solertique ejusdem cura probati & examinati, Historia. Historia Natural y Médica del Principado de Asturias. Facsímil 1998. Servicio de Publicaciones. Principado de Asturias, Oviedo, 480 pp.
  43. Dal Corso, J., Roghi, G., Ragazzi, E., Angelini, I., Giaretta, A., Soriano, C., Delclòs, X. & Jenkyns, H.C. (2013) Physico-chemical analysis of Albian (Lower Cretaceous) amber from San Just (Spain): implications for palaeoenvironmental and palaeoecological studies. Geologica Acta, 11 (3), 359–370. https:/doi.org/10.1344/105.000001871
  44. Davranoglou, L.R., Pérez-de la Fuente, R., Baňař, P. & Peñalver, E. (2024) The first unique-headed bug (Hemiptera: Enicocephalomorpha) from Cretaceous Iberian amber, and the Gondwanan connections of its palaeoentomological fauna. Papers in Palaeontology, 10 (2), e1550. https://doi.org/10.1002/spp2.1550
  45. Delclòs, X., Arillo, A., Peñalver, E., Barrón, E., Soriano, C., López Del Valle, R., Bernárdez, E., Corral, C. & Ortuño, V.M. (2007) Fossiliferous amber deposits from the Cretaceous (Albian) of Spain. Comptes Rendus Palevol, 6 (1), 135–149. https://doi.org/10.1016/j.crpv.2006.09.003
  46. Delclòs, X., Peñalver, E., Arillo, A., Engel, M.S., Nel, A., Azar, D. & Ross, A. (2016) New mantises (Insecta: Mantodea) in Cretaceous ambers from Lebanon, Spain, and Myanmar. Cretaceous Research, 60, 91–108. https://doi.org/10.1016/j.cretres.2015.11.001
  47. Dunlop, J.A., Penney, D. & Jekel, D. (2023) A summary list of fossil spiders and their relatives. In World Spider Catalog. Natural History Museum Bern, Available from: http://wsc.nmbe.ch (Accessed 15 June 2024).
  48. Engel, M.S. & Delclòs, X. (2010) Primitive termites in Cretaceous amber from Spain and Canada (Isoptera). Journal of the Kansas Entomological Society, 83 (2), 111–128. https://doi.org/10.2317/JKES0908.06.1
  49. Engel, M.S., Ortega-Blanco, J. & Azevedo, C.O. (2016) A new bethylid wasp in Lebanese Early Cretaceous amber (Hymenoptera: Chrysidoidea), with comments on other Mesozoic taxa. American Museum Novitates, 3855, 1–14. https://doi.org/10.1206/3855.1
  50. Engel, M.S., Ortega-Blanco, J. & McKellar, R.C. (2013a) New scolebythid wasps in Cretaceous amber from Spain and Canada, with implications for the phylogeny of the family (Hymenoptera: Scolebythidae). Cretaceous Research, 46, 31–42. https://doi.org/10.1016/j.cretres.2013.09.003
  51. Engel, M.S., Ortega-Blanco, J., Soriano, C., Grimaldi, D.A. & Delclòs, X. (2013b) A new lineage of enigmatic diaprioid wasps in Cretaceous Amber (Hymenoptera: Diaprioidea). American Museum Novitates, 2013, 1–23. https://doi.org/10.1206/3771.2
  52. Engel, M.S., Peris, D., Chatzimanolis, S. & Delclòs, X. (2015) An earwig (Insecta: Dermaptera) in Early Cretaceous amber from Spain. Insect Systematics & Evolution, 46, 291–300. https://doi.org/10.1163/1876312X-45032121
  53. Golub, V.B., Popov, Yu.A. & Arillo, A. (2012) Hispanocaderidae n. fam. (Hemiptera: Heteroptera: Tingoidea), one of the oldest lace bugs from the Lower Cretaceous Álava amber (Spain). Zootaxa, 3270 (1), 41–50. https://doi.org/10.11646/zootaxa.3270.1.3
  54. González-Fernández, B., Meléndez-Casares, E., Gutiérrez-Claverol, M. & García-Ramos, J.C. (2004) Litoestratigrafía del sector occidental de la cuenca cretácica de Asturias. Trabajos de Geología, Universidad de Oviedo, 24, 43–80.
  55. González-Fernández, B., Menéndez-Casares, E., Gutiérrez-Claverol, M. & García-Ramos, J.C. (2005) Discusión sobre la Revisión y síntesis litoestratigráfica del sector occidental de la cuenca cretácica de Asturias. Contrarréplica. Trabajos de Geología, Universidad de Oviedo, 25, 117–137.
  56. Grimaldi, D. & Arillo, A. (2008) The Tethepomyiidae, a new family of enigmatic Cretaceous Diptera. Alavesia, 2, 259–265.
  57. Grimaldi, D.A., Arillo, A., Cumming, J.M. & Hauser, M. (2011) Brachyceran Diptera (Insecta) in Cretaceous ambers, Part IV, Significant new orthorrhaphous taxa. ZooKeys, 148, 293–332. https://doi.org/10.3897/zookeys.148.1809
  58. Grimaldi, D.A., Cumming, J.M. & Arillo, A. (2009) Chimeromyiidae, a new family of eremoneuran Diptera from the Cretaceous. Zootaxa, 2078 (1), 34–54. https://doi.org/10.11646/zootaxa.2078.1.2
  59. Jałoszyński, P. & Peris, D. (2016) Cretaceous amber inclusions of Spain and Myanmar demonstrate early diversification and wide dispersal of Cephenniitae (Coleoptera: Staphylinidae: Scydmaeninae). Cretaceous Research, 57, 190–198. https://doi.org/10.1016/j.cretres.2015.09.002
  60. Kania, I., Krzemiński, W. & Arillo, A. (2016) First representative of the genus Helius Lepeletier and Serville, 1828 (Diptera, Limoniidae) from the Lower Cretaceous Álava amber (Spain). Cretaceous Research, 63, 33–38. https://doi.org/10.1016/j.cretres.2016.02.018
  61. Kania, I., Krzemiński, W. & Arillo, A. (2017) A new peculiar species of the genus Helius Lepeletier & Serville, 1828 (Diptera, Limoniidae) from Cretaceous Álava amber (Spain). Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107 (2-3), 1–7. https://doi.org/10.1017/S1755691017000299
  62. Kania-Kłosok, I., Arillo, A., Tuchowski, M., Zhang, Q. & Krzemiński, W. (2022) Early stage of evolution of Gonomyia (Diptera: Limoniidae), new significant discoveries in the Cretaceous Iberian and Kachin amber. Scientific Reports, 12, 21118. https://doi.org/10.1038/s41598-022-25803-0
  63. Kania-Kłosok, I., Krzemiński, W. & Arillo, A. (2021a) Two new long‑rostrum cranefly species from the Cretaceous Iberian amber (Diptera, Limoniidae, Helius). Scientific Reports, 11, 12851. https://doi.org/10.1038/s41598-021-91803-1
  64. Kania-Kłosok, I., Krzemiński, W., Kopeć, K. & Arillo, A. (2021b) The oldest evolutionary lineage of Trichoneura Loew, 1850 (Diptera, Limoniidae) and the first evidence of this genus in Cretaceous Spanish amber. Insects, 12, 411. https://doi.org/10.3390/insects12050411
  65. Krzemiński, W. & Arillo, A. (2007) Alavia neli n. gen. and n. sp.—the first Limoniidae (Diptera) from the Lower Cretaceous amber of Álava (Spain). Alavesia, 1, 11–13.
  66. Krzemiński, W., Kania-Kłosok, I., Arillo, Kopeć, K., Santos, D. & Soszyńska, A. (2024) The evolutionary history and biogeographical distribution of the Mesozoic relic genus Chilelimnophila (Diptera, Limoniidae). Zoological Journal of the Linnean Society, 202, zlae119. https://doi.org/10.1093/zoolinnean/zlae119
  67. Kvaček, J., Barrón, E., Heřmanová, Z., Mendes, M., Karch, J., Žemlička, J. & Dudák, J. (2018) Araucarian conifer from late Albian amber of northern Spain. Papers in Palaeontology, 4 (4), 643–656. https://doi.org/10.1002/spp2.1223
  68. Lepeco, A. & Melo, G.A.R. (2024) Revisiting the phylogeny of the scolebythid wasps (Hymenoptera: Aculeata) through Bayesian model evaluation and parsimony, with description of a new fossil family of Chrysidoidea. Zoological Journal of the Linnean Society, 202 (1), 57–85. https://doi.org/10.1093/zoolinnean/zlad113
  69. Li, L.F., Kopylov, D.S., Rasnitsyn, A.P., Yang, J.T., Shih, C.K. & Ren, D. (2025) New wasps of Maimetshidae (Hymenoptera: Ceraphronoidea) from the Mid-Cretaceous Myanmar amber. Insects, 16 (3), 237. https://doi.org/10.3390/insects16030237
  70. Li, S., Yoshizawa, K., Wang, Q.Z., Ren, D., Bai, M. & Yao, Y.Z. (2022) New genus and species of Empheriidae (Insecta: Psocodea: Trogiomorpha) and their implication for the phylogeny of Infraorder Atropetae. Frontiers in Ecology and Evolution, 10, 907903. https:/doi.org/10.3389/fevo.2022.907903
  71. Lozano, R.P., Pérez-de la Fuente, R., Barrón, E., Rodrigo, A., Viejo, J.L. & Peñalver, E. (2020) Phloem sap in Cretaceous ambers as abundant double emulsions preserving organic and inorganic residues. Scientific Reports, 10, 9751. https://doi.org/10.1038/s41598-020-66631-4
  72. Lukashevich, E. & Arillo, A. (2016) New Eoptychoptera (Insecta: Diptera, Ptychopteridae) from the Lower Cretaceous of Spain. Cretaceous Research, 58, 254–264. https:/doi.org/10.1016/j.cretres.2015.10.013
  73. Martín-Chivelet, J., Berástegui, X., Rosales, I., Vilas, L., Vera, J.A., Caus, E., Gräfe, K.-U., Mas, R., Puig, C., Segura, M., Robles, S., Floquet, M., Quesada, S., Ruiz-Ortiz, P.A., Fregenal-Martínez, M.A., Salas, R., Arias, C., García, A., Martín Algarra, A., Meléndez, M.N., Chacón, B., Molina, J.M., Sanz, J.L., Castro, J.M., García-Hernández, M., Carenas, B., García-Hidalgo, J., Gil, J. & Ortega, F. (2002) 12. Cretaceous. In: Gibbons, W. & Moreno, M.T. (Eds), The geology of Spain. Geological Society, London, 255–292.
  74. Martínez-Torres, L., Pujalte, V. & Robles, S. (2003) Los yacimientos de ámbar del Cretácico Inferior de Montoria-Peñacerrada (Álava, Cuenca Vasco-Cantábrica): Estratigrafía, reconstrucción paleogeográfica y estructura tectónica. Estudios del Museo de Ciencias Naturales de Álava, 18, 9–32.
  75. Mas, R., García, A., Salas, R., Meléndez, A., Alonso, A., Aurell, M., Bádenas, B., Benito, M.I., Carenas, B., García-Hidalgo, J.F., Gil, J. & Segura, M. (2004) Segunda fase de rifting: Jurásico Superior-Cretácico Inferior. In: Vera, J.A. (Ed.), Geología de España. Sociedad Geológica de España e Instituto Geológico y Minero de España, Madrid, 503–551.
  76. Menor-Salván, C., Simoneit, B.R.T., Ruiz-Bermejo M. & Alonso, J. (2016) The molecular composition of Cretaceous ambers: Identification and chemosystematic relevance of 1,6-dimethyl-5-alkyltetralins and related bisnorlabdane biomarkers. Organic Geochemistry, 93, 7–21. https://doi.org/10.1016/j.orggeochem.2015.12.010
  77. Najarro, M, Peñalver, E., Pérez-de la Fuente, R., Ortega-Blanco, J., Menor-Salván, C., Barrón, E., Soriano, C., Rosales, I., López Del Valle, R., Velasco, F., Tornos, F., Daviero-Gomez, V., Gomez, B. & Delclòs, X. (2010) Review of the El Soplao amber outcrop, Early Cretaceous of Cantabria, Spain. Acta Geologica Sinica, 84 (4), 959–976. https://doi.org/10.1111/j.1755-6724.2010.00258.x
  78. Najarro, M., Peñalver, E., Rosales, I., Pérez-de la Fuente, R., Daviero-Gomez, V., Gomez, B. & Delclòs, X. (2009) Unusual concentration of Early Albian arthropod-bearing amber in the Basque-Cantabrian Basin (El Soplao, Cantabria, Northern Spain): palaeoenvironmental and palaeobiological implications. Geologica Acta, 7, 363–387.
  79. Nel, P., Peñalver, E., Azar, D., Hodebert, G. & Nel, A. (2010) Modern thrips families Thripidae and Phlaeothripidae in Early Cretaceous amber (Insecta: Thysanoptera). Annales de la Société entomologique de France (N.S.), 46 (1-2), 154–163. https://doi.org/10.1080/00379271.2010.10697651
  80. Olmi, M., Rasnitsyn, A.P., Brothers, D.J. & Guglielmino, A. (2014) The first fossil Embolemidae (Hymenoptera: Chrysidoidea) from Burmese amber (Myanmar) and Orapa Kimberlitic deposits (Botswana) and their phylogenetic significance. Journal of Systematic Palaeontology, 12 (6), 623–635. https://doi.org/10.1080/14772019.2013.829533
  81. Ortega-Blanco, J. & Engel, M.S. (2013) Bethylidae from Early Cretaceous Spanish amber (Hymenoptera: Chrysidoidea). Journal of the Kansas Entomological Society, 86, 264–276. https://doi.org/10.2317/JKES130312.1
  82. Ortega-Blanco, J., Bennett, D.J., Delclòs, X. & Engel, M.S. (2009) A primitive aphidiine wasp in Albian amber from Spain and a Northern Hemisphere origin for the subfamily (Hymenoptera: Braconidae: Aphidiinae). Journal of the Kansas Entomological Society, 82, 273–282. https://doi.org/10.2317/JKES0812.08.1
  83. Ortega-Blanco, J., Delclòs, X. & Engel, M.S. (2011a) A protorhyssaline wasp in Early Cretaceous amber from Spain (Hymenoptera: Braconidae). Journal of the Kansas Entomological Society, 84, 51–57. https://doi.org/10.2317/JKES100728.1
  84. Ortega-Blanco, J., Delclòs, X. & Engel, M.S. (2011b) Diverse stigmaphronid wasps in Early Cretaceous amber from Spain (Hymenoptera: Ceraphronoidea: Stigmaphronidae). Cretaceous Research, 32, 762–773. https://doi.org/10.1016/j.cretres.2011.05.004
  85. Ortega-Blanco, J., Delclòs, X. & Engel, M.S. (2011c) The wasp family Embolemidae in Early Cretaceous amber from Spain (Hymenoptera: Chrysidoidea). Journal of the Kansas Entomological Society, 84, 36–42. https://doi.org/10.2317/JKES100628.1
  86. Ortega-Blanco, J., Delclòs, X., Peñalver, E. & Engel, M.S. (2011d) Serphitid wasps in early Cretaceous amber from Spain (Hymenoptera: Serphitidae). Cretaceous Research, 32, 143–154. https://doi.org/10.1016/j.cretres.2010.11.004
  87. Ortega-Blanco, J., McKellar, R.C. & Engel, M.S. (2014) Diverse scelionid wasps from Early Cretaceous Álava amber, Spain (Hymenoptera: Platygastroidea). Bulletin of Geosciences, 89, 553–571. https://doi.org/10.3140/bull.geosci.1463
  88. Ortega-Blanco, J., Peñalver, E., Delclòs, X. & Engel, M.S. (2011e) False fairy wasps in early Cretaceous amber from Spain (Hymenoptera: Mymarommatoidea). Palaeontology, 54, 511–523. https://doi.org/10.1111/j.1475-4983.2011.01049.x
  89. Ortega-Blanco, J., Rasnitsyn, A.P. & Delclòs, X. (2008) First record of anaxyelid woodwasps (Hymenoptera: Anaxyelidae) in Lower Cretaceous Spanish amber. Zootaxa, 1937 (1), 39–50. https://doi.org/10.11646/zootaxa.1937.1.3
  90. Ortega-Blanco, J., Rasnitsyn, A.P. & Delclòs, X. (2010) A new family of ceraphronoid wasps from Early Cretaceous Álava amber, Spain. Acta Palaeontologica Polonica, 55 (2), 265–276. https://doi.org/10.4202/app.2009.0014
  91. Pardo, G., Ardevol, L. & Villena, J. (1991) Mapa Geológico de España E 1:200.000, hoja nº40 (Daroca). ITGE, Madrid.
  92. Penney, D. (2006) The oldest lagonomegopid spider, a new species in Lower Cretaceous amber from Álava, Spain. Geologica Acta, 4 (3), 377–382.
  93. Penney, D. & Ortuño, V.M. (2006) Oldest true orb-weaving spider (Araneae: Araneidae). Biology Letters, 2, 447–450. https://doi.org/10.1098/rsbl.2006.0506
  94. Peñalver, E. (2011) Ámbar cretácico de San Just (Teruel): el estudio de los insectos que convivieron con los dinosaurios. Isurus, 4, 24–36.
  95. Peñalver, E. & Arillo, A. (2007) A new species of the family Hybotidae in the Lower Cretaceous amber of El Caleyu (Asturias, Spain); Alavesia prietoi n. sp. Alavesia, 1, 63–68.
  96. Peñalver, E. & Delclòs, X. (2010) Spanish amber. In: Penney, D. (Ed.), Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press, Rochdale, UK, 236–270.
  97. Peñalver, E. & Grimaldi, D.A. (2010) Latest occurrences of the Mesozoic family Elcanidae (Insecta: Orthoptera), in Cretaceous amber from Myanmar and Spain. Annales de la Société entomologique de France (N.S.), 46 (1-2), 88–99. https://doi.org/10.1080/00379271.2010.10697641
  98. Peñalver, E. & Nel, P. (2010) Hispanothrips from Early Cretaceous Spanish amber, a new genus of the resurrected family Stenurothripidae (Insecta: Thysanoptera). Annales de la Société entomologique de France (N.S.), 46 (1-2), 138–147. https://doi.org/10.1080/00379271.2010.10697649
  99. Peñalver, E. & Szwedo, J. (2010) Perforissidae (Hemiptera: Fulgoroidea) from the Lower Cretaceous San Just amber (Eastern Spain). Alavesia, 3, 97–103.
  100. Peñalver, E. & Wegierek, P. (2008) A new genus and species of the family Tajmyraphididae (Hemiptera: Sternorrhyncha) in Early Cretaceous amber from Peñacerrada I (Spain). Alavesia, 2, 187–192.
  101. Peñalver, E., Arillo, A. & Nel, A. (2022a) A review of the Cretaceous genus Eltxo (Diptera: Cecidomyiidae) with description of the new species Eltxo grimaldii from El Soplao amber. Palaeoentomology, 5 (5), 461–467. https://doi.org/10.11646/palaeoentomology.5.5.7
  102. Peñalver, E., Arillo, A., Pérez-de la Fuente, R., Riccio, M.L., Delclòs, X., Barrón, E. & Grimaldi, D.A. (2015) Long-proboscid flies as pollinators of Cretaceous gymnosperms. Current Biology, 25 (14), 1917–1923. https://doi.org/10.1016/j.cub.2015.05.062
  103. Peñalver, E., Delclòs, X. & Soriano, C. (2007) A new rich amber outcrop with palaeobiological inclusions in the Lower Cretaceous of Spain. Cretaceous Research, 28, 791–802. https://doi.org/10.1016/j.cretres.2006.12.004
  104. Peñalver, E., González-Fernández, B., López Del Valle, R., Barrón, E., Lozano, R.P., Rodrigo, A., Pérez-de la Fuente, R., Menéndez-Casares, E. & Sarto i Monteys, V. (2018) Un nuevo yacimiento de ámbar cretácico en Asturias (norte de España): resultados preliminares de la excavación paleontológica de 2017 en La Rodada (La Manjoya). Cuadernos del Museo Geominero, 27, 289–299.
  105. Peñalver, E., Grimaldi, D.A. & Delclòs, X. (2006) Early Cretaceous spider web with its prey. Science, 312, 1761. https://doi.org/10.1126/science.1126628
  106. Peñalver, E., Labandeira, C.C., Barrón, E., Delclòs, X., Nel, P., Nel, A., Tafforeau, P. & Soriano, C. (2012) Thrips pollination of Mesozoic gymnosperms. Proceedings of the National Academy of Sciences, 109 (22), 8623–8628. https://doi.org/10.1073/pnas.1120499109
  107. Peñalver, E., Nel, A. & Nel, P. (2022b) An apterous Tubulifera (Insecta, Thysanoptera, Phlaeothripidae) preserved in Spanish Cretaceous amber. Historical Biology, 34 (2), 1–7. https://doi.org/10.1080/08912963.2021.2023139
  108. Peñalver, E., Ortega-Blanco, J., Nel, A. & Delclòs, X. (2010) Mesozoic Evaniidae (Insecta: Hymenoptera) in Spanish amber: Reanalysis of the phylogeny of the Evanioidea. Acta Geologica Sinica, 84, 809–827. https://doi.org/10.1111/j.1755-6724.2010.00257.x
  109. Peñalver, E., Peña-Kairath, C., Barrón, E., Nel, P., Nel, A., Delclòs, X., Peris, D., Solórzano-Kraemer, M.M. & Rodrigo, A. (2025a) Diverse Mesozoic thrips carrying pollen during the gymnosperm-to-angiosperm plant-host ecological shift. iScience, 28, 112108. https://doi.org/10.1016/j.isci.2025.112108
  110. Peñalver, E., Speranza, M. & Delclòs, X. (2025b) Fungal cortices in kauri resin from New Zealand as taphonomic and ecological analogues of Cretaceous amber cortices: an actualistic approach. Spanish Journal of Palaeontology, 40 (1), 133–146. https://doi.org/10.7203/sjp.29855
  111. Pérez-de la Fuente, R. & Peñalver, E. (2019) A mantidfly in Cretaceous Spanish amber provides insights into the evolution of integumentary specialisations on the raptorial foreleg. Scientific Reports, 9, 13248. https://doi.org/10.1038/s41598-019-49398-1
  112. Pérez-de la Fuente, R., Delclòs, X., Peñalver, E. & Arillo, A. (2011) Biting midges (Diptera: Ceratopogonidae) from the Early Cretaceous El Soplao amber (N Spain). Cretaceous Research, 32, 750–761. https://doi.org/10.1016/j.cretres.2011.05.003
  113. Pérez-de la Fuente, R., Delclòs, X., Peñalver, E. & Engel, M.S. (2019) A new dustywing (Neuroptera: Coniopterygidae) from the Early Cretaceous amber of Spain. Palaeoentomology, 2 (3), 279–288. https://doi.org/10.11646/palaeoentomology.2.3.13
  114. Pérez-de la Fuente, R., Delclòs, X., Peñalver, E., Speranza, M., Wierzchos, J., Ascaso, C. & Engel, M.S. (2012a) Early evolution and ecology of camouflage in insects. Proceedings of the National Academy of Sciences, 109 (52), 1414–1419. https://doi.org/10.1073/pnas.1213775110
  115. Pérez-de la Fuente, R., Nel, A., Peñalver, E. & Delclòs, X. (2010) A new Early Cretaceous snakefly (Raphidioptera: Mesoraphidiidae) from El Soplao amber (Spain). Annales de la Société Entomologique de France (N.S.), 46 (1-2), 108–115. https://doi.org/10.1080/00379271.2010.10697644
  116. Pérez-de la Fuente, R., Peñalver, E., Delclòs, X. & Engel, M.S. (2012b) Snakefly diversity in Early Cretaceous amber from Spain (Neuropterida, Raphidioptera). ZooKeys, 204, 1–40. https://doi.org/10.3897/zookeys.204.2740
  117. Pérez-de la Fuente, R., Peñalver, E. & Engel, M.S. (2021) Beaded lacewings (Neuroptera: Berothidae) in amber from the Lower Cretaceous of Spain. Cretaceous Research, 119, 104705. https://doi.org/10.1016/j.cretres.2020.104705
  118. Pérez-de la Fuente, R., Peñalver, E. & Ortega-Blanco, J. (2012c) A new species of the diverse Cretaceous genus Cretevania Rasnitsyn, 1975 (Hymenoptera: Evaniidae) from Spanish amber. Zootaxa, 3514 (1), 70–78. https://doi.org/10.11646/zootaxa.3514.1.4
  119. Pérez-de la Fuente, R., Saupe, E.E. & Selden, P.A. (2013) New lagonomegopid spiders (Araneae: †Lagonomegopidae) from Early Cretaceous Spanish amber. Journal of Systematic Palaeontology, 11 (5), 531–553. https://doi.org/10.1080/14772019.2012.725679
  120. Peris, D. (2017) Early Cretaceous origin of pollen-feeding beetles (Insecta: Coleoptera: Oedemeridae). Cladistics, 33, 268–278. https://doi.org/10.1111/cla.12168
  121. Peris, D. & Delclòs, X. (2015) Fossil Monotomidae (Coleoptera: Polyphaga) from Laurasian Cretaceous amber. Organisms Diversity & Evolution, 15, 1–10. https://doi.org/10.1007/s13127-015-0205-y
  122. Peris, D. & Fanti, F. (2018) Molliberus albae gen. et sp. nov., the oldest Laurasian soldier beetle (Coleoptera: Cantharidae) from the Lower Cretaceous Spanish amber. Cretaceous Research, 91, 263–268. https://doi.org/10.1016/j.cretres.2018.07.003
  123. Peris, D. & Ruzzier, E. (2013) A new tribe, new genus, and new species of Mordellidae (Coleoptera: Tenebrionoidea) from the Early Cretaceous amber of Spain. Cretaceous Research, 45, 1–6. https://doi.org/10.1016/j.cretres.2013.07.002
  124. Peris, D., Chatzimanolis, S. & Delclòs, X. (2014a) Diversity of rove beetles (Coleoptera: Staphylinidae) in Early Cretaceous Spanish amber. Cretaceous Research, 48, 85–95. https://doi.org/10.1016/j.cretres.2013.11.008
  125. Peris, D., Davis, S.R., Engel, M.S. & Delclòs, X. (2014b) An evolutionary history embedded in amber: reflection of the Mesozoic shift in weevil-dominated (Coleoptera: Curculionoidea) faunas. Zoological Journal of the Linnean Society, 171, 534–553. https://doi.org/10.1111/zoj.12149
  126. Peris, D., Lyubarsky, G.Yu & Perkovsky, E.E. (2017a) A new genus of silken-fungus beetle (Coleoptera: Cryptophagidae) from the Spanish Cretaceous amber. Cretaceous Research, 78, 191–195. https://doi.org/10.1016/j.cretres.2017.06.018
  127. Peris, D., Maier, C.A., Sánchez-García, A. & Delclòs, X. (2015a) The oldest known riffle beetle (Coleoptera: Elmidae) from Early Cretaceous Spanish amber. Comptes Rendus Palevol, 14, 181–186. https://doi.org/10.1016/j.crpv.2014.11.005
  128. Peris, D., Peñalver, E., Delclòs, X., Barrón, E., Pérez-de la Fuente, R. & Labandeira, C.C. (2017b) False blister beetles and the expansion of gymnosperm–insect pollination modes before angiosperm dominance. Current Biology, 27 (6), 897–904. https://doi.org/10.1016/j.cub.2017.02.009
  129. Peris, D., Philips, T.K. & Delclòs, X. (2015b) Ptinid beetles from the Cretaceous gymnosperm-dominated forests. Cretaceous Research, 52, 440–452. https://doi.org/10.1016/j.cretres.2014.02.009
  130. Perreau, M. (2019) Cretaciella sorianoae gen. et sp. nov. (Coleoptera, Leiodidae, Cholevinae, Oritocatopini), anophthalmic species from Albian amber of the Escucha Formation (Alava, Spain). Geosciences, 9 (12), 521. https://doi.org/10.3390/geosciences9120521
  131. Perrichot, V. (2009) Long-tailed wasps (Hymenoptera: Megalyridae) from Cretaceous and Paleogene European amber. Paleontological Contributions, 1, 1–35. https:/doi.org/10.17161/PC.1808.5468
  132. Perrichot, V., Ortega-Blanco, J., McKellar, R.C., Delclòs, X., Azar, D., Nel, A., Tafforeau, P. & Engel, M.S. (2011) New and revised maimetshid wasps from Cretaceous ambers (Hymenoptera, Maimetshidae). ZooKeys, 130, 421–453. https://doi.org/10.3897/zookeys.130.1453
  133. Peyrot, D., Rodríguez-López, J.P., Lassaletta, L., Meléndez, N. & Barrón, E. (2007) Contributions to the palaeoenvironmental knowledge of the Escucha Formation in the Lower Cretaceous Oliete Sub-basin, Teruel, Spain. Comptes Rendus Palevol, 6, 469–481. https://doi.org/10.1016/j.crpv.2007.09.014
  134. Querol, X., Salas, R., Pardo, G. & Ardévol, L. (1992) Albian coal-bearing deposits of the Iberian Range in northeaster Spain. In: McCabe, P.J. & Parrish, J.T. (Eds), Controls on the distribution and quality of Cretaceous coals. The Geologial Society of America, Special paper 267, 193–208. https://doi.org/10.1130/SPE267-p193
  135. Rasnitsyn, A.P., Maalouf, M., Maalouf, R. & Azar, D. (2022) New Serphitidae and Gallorommatidae (Insecta: Hymenoptera: Microprocta) in the Early Cretaceous Lebanese amber. Palaeoentomology, 5 (2), 120–136. https://doi.org/10.11646/palaeoentomology.5.2.4
  136. Robles, S., Aranburu, A. & Apraiz, A. (2014) La Cuenca Vasco-Cantábrica: génesis y evolución tectonosedimentaria. Enseñanza de las Ciencias de la Tierra, 22.2, 99–114.
  137. Rodríguez-López, J.P., Meléndez, N., Soria, A.R. & De Boer, P. (2009) Reinterpretación estratigráfica y sedimentológica de las formaciones Escucha y Utrillas de la Cordillera Ibérica. Revista de la Sociedad Geológica de España, 22, 163–219.
  138. Rodrigo, A., Peñalver, E., López Del Valle, R., Barrón, E. & Delclòs, X. (2018) The heritage interest of the Cretaceous amber outcrops in the Iberian Peninsula, and their management and protection. Geoheritage, 10, 511–523. https://doi.org/10.1007/s12371-018-0292-1
  139. Salas, R., Guimerà, J., Bover-Arnal, T. & Nebot, M. (2019) The Iberian-Catalan linkage: the Maestrat and Garraf basins. In: Quesada, C. & Oliveira, J.T. (Eds), The geology of Iberia: a geodynamic approach. Springer, Volume 3: The Alpine Cycle, 228–230.
  140. Sánchez-García, A. & Engel, M.S. (2016a) Springtails from the Early Cretaceous amber of Spain (Collembola: Entomobryomorpha), with an annotated checklist of fossil Collembola. American Museum Novitates, 3862, 1–47. https://doi.org/10.1206/3862.1
  141. Sánchez-García, A. & Engel, M.S. (2016b) Long-term stasis in a diverse fauna of Early Cretaceous springtails (Collembola: Symphypleona). Journal of Systematic Palaeontology, 15 (7), 513–537. https://doi.org/10.1080/14772019.2016.1194575
  142. Sánchez-García, A., Arillo, A. & Nel, A. (2016) The first water measurers from the Lower Cretaceous amber of Spain (Heteroptera, Hydrometridae, Heterocleptinae). Cretaceous Research, 57, 111–121. https://doi.org/10.1016/j.cretres.2015.07.010
  143. Sánchez-García, A., Nel, A., Arillo, A. & Solórzano-Kraemer, M.M. (2017) The semi-aquatic pondweed bugs of a Cretaceous swamp. PeerJ, 5, e3760. https://doi.org/10.7717/peerj.3760
  144. Sánchez-García, A., Palencia, L., Delclòs, X. & Peñalver, E. (2024) False scorpions (Arachnida: Pseudoscorpiones) from Lower Cretaceous Spanish amber. Papers in Palaeontology, 10 (6), e1608. https://doi.org/10.1002/spp2.1608
  145. Sánchez-García, A., Peñalver, E., Delclòs, X. & Engel, M.S. (2020) Early Cretaceous termites in amber from northern Spain (Isoptera). Cretaceous Research, 110, 104385. https://doi.org/10.1016/j.cretres.2020.104385
  146. Sánchez-García, A., Peñalver, E., Delclòs, X. & Engel, M.S. (2021) Terrestrial isopods from Spanish amber (Crustacea: Oniscidea): Insights into the Cretaceous soil biota. American Museum Novitates, 3974, 1–32. https://doi.org/10.1206/3974.1
  147. Sánchez-García, A., Peñalver, E., Pérez-de la Fuente, R. & Delclòs, X. (2014) A rich and diverse tanaidomorphan (Crustacea: Tanaidacea) assemblage associated with Early Cretaceous resin-producing forests in North Iberia: palaeobiological implications. Journal of Systematic Palaeontology, 13 (8), 645–676. https://doi.org/10.1080/14772019.2014.944946
  148. Santer, M., Álvarez-Parra, S., Nel, A., Peñalver, E. & Delclòs, X. (2022) New insights into the enigmatic Cretaceous family Spathiopterygidae (Hymenoptera: Diaprioidea). Cretaceous Research, 133, 105128. https://doi.org/10.1016/j.cretres.2021.105128
  149. Saupe, E.E., Pérez-de la Fuente, R., Selden, P.A., Delclòs, X., Tafforeau, P. & Soriano, C. (2012) New Orchestina Simon, 1882 (Araneae: Oonopidae) from Cretaceous ambers of Spain and France: first spiders described using phase-contrast X-ray synchrotron microtomography. Palaeontology, 55, 127–143. https://doi.org/10.1111/j.1475-4983.2011.01123.x
  150. Sidorchuk, E.A. & Norton, R.A. (2011) The fossil mite family Archaeorchestidae (Acari, Oribatida) I: redescription of Strieremaeus illibatus and synonymy of Strieremaeus with Archaeorchestes. Zootaxa, 2993 (1), 34–58. https://doi.org/10.11646/zootaxa.2993.1.3
  151. Simón-Benito, J.C., Ortuño, V.M. & Espantaleón, D. (2002) Colémbolos (Collembola, Insecta) del ámbar Cretácico de Álava (Cuenca Vasco-Cantábrica, Norte de España). Estudios del Museo de Ciencias Naturales de Álava, 17, 83–92.
  152. Sinclair, B.J. & Kirk-Spriggs, A.H. (2010) Alavesia Waters and Arillo—a Cretaceous-era genus discovered extant on the Brandberg Massif, Namibia (Diptera: Atelestidae). Systematic Entomology, 35, 268–276. https://doi.org/10.1111/j.1365-3113.2009.00506.x
  153. Skibińska, K. & Arillo, A. (2017) The first Tanyderidae (Diptera) from Lower Cretaceous Álava amber (Spain). Historical Biology, 31 (4), 1–7. https:/doi.org/10.1080/08912963.2017.1398747
  154. Solórzano-Kraemer, M.M., Delclòs, X., Clapham, M., Arillo, A., Peris, D., Jäger, P., Stebner, F. & Peñalver, E. (2018) Arthropods in modern resins reveal if amber accurately recorded forest arthropod communities. Proceedings of the National Academy of Sciences, 115 (26), 6739–6744. https://doi.org/10.1073/pnas.1802138115
  155. Solórzano-Kraemer, M.M., Sinclair, B.J., Arillo, A. & Álvarez-Parra, S. (2023) A new genus of dance fly (Diptera: Empidoidea: Hybotidae) from Cretaceous Spanish ambers and introduction to the fossiliferous amber outcrop of La Hoya (Castellón Province, Spain). PeerJ, 11, e14692. https://doi.org/10.7717/peerj.14692
  156. Soszyńska-Maj, A., Krzemińska, E., Pérez-de la Fuente, R., Wang, J.S., Szpila, K., Skibińska, K., Kopeć, K. & Krzemiński, W. (2022) Evolution of sexual conflict in scorpionflies. eLife, 11, e70508. https://doi.org/10.7554/eLife.70508
  157. Speranza, M., Ascaso, C., Delclòs, X. & Peñalver, E. (2015) Cretaceous mycelia preserving fungal polysaccharides: Taphonomic and paleoecological potential of microorganisms preserved in fossil resins. Geologica Acta, 13 (4), 363–385. https:/doi.org/10.1344/geologicaacta2015.13.4.8
  158. Szadziewski, R. & Arillo, A. (1998) Biting midges (Diptera: Ceratopogonidae) from the Lower Cretaceous amber from Alava, Spain. Polskie Pismo Entomologiczne, 67, 291–298.
  159. Szadziewski, R. & Arillo, A. (2003) The oldest fossil record of the extant subgenus Leptoconops (Leptoconops) (Diptera: Ceratopogonidae). Acta Zoologica Cracoviensia, 46 (suppl), 271–275.
  160. Szadziewski, R., Arillo, A., Urbanek, A. & Sontag, E. (2016) Biting midges of the extinct genus Protoculicoides Boesel from Lower Cretaceous amber of San Just, Spain and new synonymy in recently described fossil genera (Diptera: Ceratopogonidae). Cretaceous Research, 58, 1–9. https://doi.org/10.1016/j.cretres.2015.09.016
  161. Talamas, E.J., Johnson, N.F., Shih, C. & Ren, D. (2019) Proterosceliopsidae: A new family of Platygastroidea from Cretaceous amber. Journal of Hymenoptera Research, 73, 3–38. https://doi.org/10.3897/jhr.73.32256
  162. Ulitzka, M.R. (2022) New genera and species of Rohrthripidae (Thysanoptera: Tubulifera) from Burmese Cretaceous amber. Zootaxa, 5162 (1), 1–36. https://doi.org/10.11646/zootaxa.5162.1.1
  163. Vilanova y Piera, J. (1860) Manual de geología aplicada a la agricultura y a las artes industriales. Imprenta Nacional, Madrid, 1, 47+XIX+384 pp.
  164. Vonk, R. & Schram, F.R. (2007) Three new tanaid species (Crustacea, Peracarida, Tanaidacea) from the Lower Cretaceous Álava amber in northern Spain. Journal of Paleontology, 81 (6), 1502–1509. https://doi.org/10.1666/05-020.1
  165. Waters, S.B. & Arillo, A. (1999) A new genus of Hybotidae (Diptera, Empidoidea) from Lower Cretaceous amber of Álava (Spain). Studia dipterologica, 6 (1), 59–66.
  166. Wegierek, P., Cai, C.Y. & Huang, D.Y. (2019) New genera and species of aphids (Hemiptera, Aphidomorpha, Burmitaphididae) in mid-Cretaceous Burmese amber. Cretaceous Research, 93, 318–329. https://doi.org/10.1016/j.cretres.2018.07.016
  167. Wunderlich, J. (2015) On the evolution and the classification of spiders, the Mesozoic spider faunas, and descriptions of new Cretaceous taxa mainly in amber from Myanmar (Burma) (Arachnida: Araneae). Beiträge zur Araneologie, 9, 21–408.