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Published: 2022-12-23

The first Archijassidae from the Middle Triassic of China (Hemiptera, Cicadomorpha, Membracoidea)

State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
State Key Laboratory of Palaeobiology and Stratigraphy, Center for Excellence in Life and Paleoenvironment, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China
Hemiptera Cicadomorpha Membracoidea Karajassinae Tongchuan entomofauna taxonomy Eocicadellium new genus


Eocicadellium grimaldii gen. et sp. nov., the oldest representative and first Triassic member of the subfamily Karajassinae Shcherbakov, 1992, is described on the basis of six forewings from the Middle-Upper Triassic Yanchang Formation of northern China, widening the duration and biogeographic distribution of this extinct subfamily. The new data represents the first known definite record of Karajassinae in China, and further explores the palaeodiversity of Triassic archijassids.


  1. Ansorge, J. (1996) Insekten aus dem oberen Lias von Grimmen (Vorpommern, Norddeutschland). Neue Paläontologische Abhandlungen, 2, 1–132.
  2. Becker-Migdisova, E.E. (1962) Nekotorye novye poluzhestkokrylye i senoedy. [Some new Hemiptera and Psocoptera]. Paelontologicheskii Zhurnal, 1, 189–104. [In Russian]
  3. Bourgoin, T., Wang, R., Asche, M., Hoch, H., Soulier-Perkins, A., Stroiński, A., Yap, S. & Szwedo, J. (2015) From micropterism to hyperpterism: recognition strategy and standardized homology-driven terminology of the forewing venation patterns in planthoppers (Hemiptera: Fulgoromorpha). Zoomorphology, 134, 63–77.
  4. Chen, J., Wang, B., Zheng, Y., Jiang, H., Jiang, T., Wang, X.L. & Zhang, H.C. (2020) The youngest record of the leafhopper family Archijassidae in Kachin amber from the lowermost Upper Cretaceous of northern Myanmar (Cicadomorpha, Cicadelloidea). Cretaceous Research, 106, 104252.
  5. Evans, J.W. (1956) Palaeozoic and Mesozoic Hemiptera (Insecta). Australian Journal of Zoology, 4, 165–158.
  6. Fu, Y.Z., Azar, D. & Huang, D.Y. (2021) The first Dysmorphoptilidae from the Middle Triassic of China (Hemiptera: Cicadomorpha). Historical Biology, 33, 3506–3512.
  7. Fu, Y.Z., Gao, J. & Huang, D.Y. (2022) Revision of the genus Sinogranulus (Hemiptera, Granulidae) with description of a new species from the Middle Triassic of China. Palaeoentomology, 5 (1), 81–89.
  8. Fu, Y.Z. & Huang, D.Y. (2022) The first maguviopseids (Hemiptera, Cicadomorpha, Prosboloidea) from the Triassic of China. Palaeoentomology, 5 (1), 76–80.
  9. Handlirsch, B.A. (1906–1908) Die fossilen Insekten und die Phylogenie der Rezenten Formen. Ein Handbuch für Paläontologen und Zoologen. Engelmann, Leipzig, 1430 pp.
  10. Hong, Y.C. (1980) Granulidae, a new family of Homoptera from the Middle Triassic of Tongchuan, Shaanxi Province. Acta Zootaxonomica Sinica, 5, 63–70. [In Chinese with English abstract]
  11. Hong, Y.C. (1984) Curvicubitidae fam. nov. (Lepidoptera, Insecta) from Middle Triassic of Shaanxi. Acta Palaeontologica Sinica, 23, 782–785. [In Chinese with English abstract]
  12. Jell, P.A. (2004) The fossil insects of Australia. Memoirs of the Queensland Museum, 50 (1), 1–124.
  13. Lambkin, K.J. (2020) Revision of Mesojassus Tillyard, 1916, from the Late Triassic of Queensland (Hemiptera: Cicadomorpha: Membracoidea: Archijassidae). Zootaxa, 4718 (3), 413–422.
  14. Lian, X.N., Cai, C.Y. & Huang, D.Y. (2022) New species of permochoristids (Insecta, Mecoptera) from the late Middle Triassic Tongchuan entomofauna in Shaanxi Province, northwestern China. Historical Biology. [Published online 07 October 2022]
  16. Liu, Z.J., Liu, S.T. & Hong, Y.C. (1985) Discovery and studying of the Triassic fauna and flora from the Niangniangmiao in Longxian, Shaanxi. Bulletin of the Xi’An Institute of Geology and Mineral Resources, 10, 105–120. [In Chinese with English abstract]
  17. Martynov, A. (1926) Jurassic fossil insects from Turkestan. 6. Homoptera and Psocoptera. Bulletin de l’Académie des Sciences de l’URSS, 20, 1349–1366.
  18. Nel, A., Prokop, J., Nel, P., Grandcolas, P., Huang, D.Y, Roques, P., Guilbert, E., Dostál, O. & Szwedo, J. (2012) Traits and evolution of wing venation pattern in paraneopteran insects. Journal of Morphology, 273, 480–506.
  19. Shcherbakov, D.E. (1992) The earliest leafhoppers (Hemiptera: Karajassidae n. fam. from the Jurassic of Karatau). Neues Jahrbuch für Geologie und Palaontologie Monatshefte, 1 (1), 39–51.
  20. Shcherbakov, D.E. (2011) New and little-known families of Hemiptera Cicadomorpha from the Triassic of Central Asia—early analogs of treehoppers and planthoppers. Zootaxa, 2836 (1), 1–26.
  21. Shcherbakov, D.E. (2012) More on Mesozoic Membracoidea (Homoptera). Russian Entomological journal, 21 (1), 15–22.
  22. Szwedo, J. (2018) The unity, diversity and conformity of bugs (Hemiptera) through time. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 107, 109–128.
  23. Tillyard, R.J. (1916) Mesozoic and Tertiary insects of Queensland and New South Wales. Geological Survey of Queensland Publication, 253, 11–64.
  24. Westwood J.O. (1854) Contributions to fossil entomology. The Quarterly Journal of the Geological Society of London, 10, 378–396.
  25. Zhang, Q.Q., Chen, J. & Zhang, H.C. (2022a) New Granulidae (Hemiptera: Scytinopteroidea) from the Middle Triassic Tongchuan Formation of NW China. Historical Biology, 34, 152–157.
  26. Zhang, Q.Q., Zheng, D.R., Wang, B. & Zhang, H.C. (2022b) A review of Triassic insects in China. Geological Society, London, Special Publications, 521 (1), 45.