Abstract
A new pachylaelapid mite species of the genus Onchodellus, O. masani sp. nov., is described and illustrated based on female specimens from Iran. The new species is compared with congeneric species with shortened peritremes. The new regional distributions for three examined species, Onchodellus hispani (Berlese), O. karawaiewi (Berlese), and O. strigifer (Berlese) are presented, and the situation of other species of the genus in Iran is revised. The current knowledge of the pachylaelapid species of Iran is discussed and a new list of the described and recorded species of the family is provided.
References
Abolghasemi, S. & Kazemi, Sh. (2016) A new record of the genus Dendrolaelaps Halbert (Acari: Mesostigmata: Digamasellidae) from Iran. In: Talaei-Hassanloui, R. (Ed.), Proceedings of the 22nd Iranian Plant Protection Congress. College of Agriculture and Natural Resources, University of Tehran, Karaj, p. 510.
Abutaleb Kermani, R., Ahadiyat, A. & Joharchi, O. (2017) A research on the distribution of soil inhabiting mites of the superfamily Eviphidoidea (Acari: Mesostigmata) in the Kashan region, Isfahan Province, Iran. In: Hajiqanbar, H. & Saboori, A. (Eds.), Program & Abstract Book of the Third International Persian Congress of Acarology. University of Tehran, Tehran, pp. 2–3.
Ahadiyat, A. & Cheraghali, Z. (2012) Faunistic study of pachylaelapid mites (Mesostigmata: Eviphidoidea: Pachylaelapidae) in Roodbaar-Ghasraan region (Shemiraanaat county) of Tehran province. In: Sarafrazi, A., Asef, M.R., Mozhdehi, M., Mozhdehi, M., Solhjouy Fard, S. & Abdollahi, T. (Eds.), Proceedings of the 20th Iranian Plant Protection Congress. Shiraz University, Shiraz, p. 428.
Ahadiyat, A., Ghasemi Moghadam, S. & Cheraghali, Z. (2016) Pachyseius masanisimilis (Mesostigmata: Eviphidoidea: Pachylaelapidae), a new species of edaphic mite from Iran. Persian Journal of Acarology, 5 (2), 109–120.
https://doi.org/10.22073/pja.v5i2.20513
Ahadiyat, A., Mašán, P., Cheraghali, Z. & Joharchi, O. (2014) First report of the subgenus Pachylaelaps (Longipachylaelaps) Mašán (Mesostigmata: Eviphidoidea: Pachylaelapidae) from Iran. Persian Journal of Acarology, 3 (1), 99–102.
https://doi.org/10.22073/pja.v3i1
Ahangaran, Y., Afshari, A., Saboori, A. & Kazemi, Sh. (2012) The fauna of the edaphic and dung dweller mites of the superfamily Eviphidoidea (Acari: Mesostigmata) in Nowshahr county, northern Iran. Taxonomy and Biosystematics, 12, 1–16. [in Persian with English abstract]
Ahangaran, Y., Afshari, A., Saboori, A., Kazemi, Sh. & Asadeh, Gh. (2010) Introduction of some pachylaelapid mites (Acari: Mesostigmata) in Nowshahr Region. In: Manzari, S. (Ed.), Proceeding of the 19th Iranian Plant Protection Congress, Vol. 1:Pests. Iranian Research Institute of Plant Protection, Tehran, p. 368.
Ahmadi, K. (2002) New records of six mite species of the genus Pachylaelaps from (Acari: Pachylaelapidae) Kerman province, Iran. In: Proceeding of the 15th Iranian Plant Protection Congress. Vol. 1. Pests. Razi University, Kermanshah, p. 149.
Alizadeh, S. & Shirdel, D. (2012) Some mesostigmatic mites (Acari: Mesostigmata) in apple orchards of Salmas region, West Azarbaijan Province, Iran. In: Sarafrazi, A., Asef, M.R., Mozhdehi, M., Mozhdehi, M., Solhjouy Fard, S. & Abdollahi, T. (Eds.), Proceedings of the 20th Iranian Plant Protection Congress. Shiraz University, Shiraz, p. 497.
Arjomandi, E. & Kazemi, Sh. (2014) Edaphic mesostigmatic mites (Acari: Mesostigmata) fauna in parks and landscapes of Kerman City. In: Ashrafjou, M. (Ed.), 3rd Integrated Pest Management Conference (IPMC). Shahid Bahonar University of Kerman, Kerman, pp. 355–363. [in Persian with English abstract]
Athias-Henriot, C. (1971) La divergence néotaxique des Gamasides (Arachnides). Bulletin Scientifique de Bourgogne, 28, 93–106.
Athias-Henriot, C. (1975) Nouvelles notes sur les Amblyseiini. II. Le relevé organotaxique de la face dorsale adulte (Gamasides, protoadéniques, Phytoseiidae). Acarologia, 17 (1), 20–29.
Babaeian, E. & Kazemi, Sh. (2011) Mites of the family Pachylaelapidae (Mesostigmata: Eviphidoidea) in Shahrekord region, Iran. In: Kazemi, Sh. & Saboori, A. (Eds.), Abstract and Proceeding Book of the First Persian Congress of Acarology. International Center for Science, High Technology & Environmental Sciences, Kerman, p. 43.
Babaeian, E., Mašán, P. & Saboori, A. (2016a) The genus Pachyseius Berlese, 1910 in Iran (Acari: Pachylaelapidae). Zootaxa, 4088 (3), 420–428.
https://doi.org/10.11646/zootaxa.4088.3.7
Babaeian, E., Mašán, P. & Saboori, A. (2016b) A new species of the genus Pachyseius Berlese (Acari: Pachylaelapidae) from Iran, with remarks on the world fauna. Persian Journal of Acarology, 5 (4), 299–309.
https://doi.org/10.22073/pja.v5i4.23553
Baroozeh, S. & Ahadiyat, A. (2012) Species composition of mites of the superfamily Eviphidoidea (Acari: Mesostigmata) in main climate zones of Tehran province, Iran. In: Murariu, D., Adam, C., Chişamera, G., Iorgu, E., Ovidiu Popa, L. & Paula Popa, O. (Eds.), Book of Abstracts of Fourth Annual Zoological Congress of “Grigore Antipa” Museum (CZGA 2012). “Ion Heliade Rădulescu” Amphitheatre of the Romanian Academy, Bucharest, p. 134.
Berlese, A. (1888) Acari Austro-Americani quos collegit Aloysius Balzan. Manipulus primus. Species novas circiter quinquaginta complectens. Bolletino della Società Entomologica Italiana, 20, 171–222, pls. 5–13.
Berlese, A. (1892) Acari, Myriopoda et Scorpiones, hucusque in Italia reperta, 64, 1–15 (text), 10 pls.
Berlese, A. (1904) Illustrazione iconografica degli Acari mirmecofili. Redia, 1, 299–474, pls. VII–XX.
Berlese, A. (1908) Elenco di generi e specie nuove di acari. Redia, 5, 1–15.
Berlese, A. (1910) Lista di nuove specie e nuovi generi di Acari. Redia, 6, 242–271.
Berlese, A. (1913) Acarotheca Italica. Fasciculi Ius et IIus. M. Ricci, Firenze, 221 pp.
Berlese, A. (1916) Centuria prima di Acari nuovi. Redia, 12, 19–67.
Berlese, A. (1920a) Acari, Myriopoda et Pseudoscorpiones hucusque in Italia reperta. Redia, 14, 77–105.
Berlese, A. (1920b) Centuria quinta di Acari nuovi. Redia, 14, 143–195.
Canestrini, G. (1881) Osservazioni intorno al genere Gamasus. Atti del Reale Istituto Veneto di Scienze, Lettere ed Arti, 7 (5), 511–525.
Costa, M. (1961) Mites recovered from the nests of the Levant Vole (Microtus guentheri) in Israel. Annals and Magazine of Natural History, Series 13, 4 (41), 257–282.
https://doi.org/10.1080/00222936108651107
Costa, M. (1963) The mesostigmatic mites associated with Copris hispanus (L.) (Coleoptera, Scarabaeidae) in Israel. Journal of the Linnaean Society, Zoology, 45 (303), 25–45.
https://doi.org/10.1111/j.1096-3642.1963.tb00485.x
Costa, M. (1966) Descriptions of the juvenile stages of Pachylaelaps hispani Berlese (Acari: Mesostigmata). Acarologia, 8 (1), 9–22.
Costa, M. (1971) Mites of the genus Pachylaelaps Berlese (Acari: Mesostigmata, Pachylaelapidae) from litter in Israel. Israel Journal of Zoology, 20 (4), 253–277.
https://doi.org/10.1080/00212210.1971.10688338
Darvishzadeh, I. & Kamali, K. (2009) Faunistic survey of mite (Acari) associated with grapevine yards in Safiabad, Khuzestan, Iran. Journal of Entomological Research, 1 (1), 79–93. [in Persian with English abstract]
Evans, G.O. & Till, W.M. (1979) Mesostigmatic mites of Britain and Ireland (Chelicerata: Acari-Parasitiformes). An introduction to their external morphology and classification. Transactions of the Zoological Society of London, 35 (2), 145–270.
https://doi.org/10.1111/j.1096-3642.1979.tb00059.x
Gholami, H. & Ostovan, H. (2013) The first report of Pachyseius humeralis (Acari: Mesostigmata: Pachylaelapidae) from Iran. Plant Protection Journal, 5, 109–113. [in Persian with English abstract]
Goncharova, A.A. & Koroleva, E.V. (1974) Two new species of the genus Pachylaelaps (Parasitiformes, Pachylaelaptidae). Zoologicheskii Zhurnal, 53, 1257–1260.
Granpayeh, S. & Ostovan, H. (2014) Mites recorded from insectary cultures in the Shiraz region of Iran. International Research Journal of Applied and Basic Sciences, 8 (5), 612–616.
Gwiazdowicz, D.J. & Gulvik, M.E. (2005) Checklist of Norwegian mesostigmatid mites (Acari, Mesostigmata). Norwegian Journal of Entomology, 52, 117–125.
Haddad Irani-Nejad, K., Hajiganbar, H.R. & Talebi Chaichi, P. (2003) Introduction of some mesostigmatic mites of sugarbeet fields in Miandoab plain. Journal of Agricultural Sciences and Natural Resources, 10 (2), 147–157. [in Persian with English abstract]
Halbert, J.N. (1915) Clare Island Survey. 39. Acarinida. Section II. – Terrestrial and Marine Acarina. Proceedings of the Royal Irish Academy, 31 (Section 2), 45–136, pls. IV–VIII.
Hamidi, P., Ostovan, H., Kamali, K. & Ahadiyat, A. (2013) Fauna of mites associated with ornamental plants in Tehran, Iran. In: Joharchi, O. & Saboori, A. (Eds.), Program & Abstract Book of the Second International Persian Congress of Acarology. Faculty of Agriculture, University of Tehran, Karaj, p. 14.
Hasanvand, I., Jafari, Sh., Kazemi, Sh. & Shakarami, J. (2015) Fauna and species diversity of edaphic mesostigmatic mites of superfamilies Eviphidoidea and Ascoidea (Acari: Mesostigmata) in Khorramabad County, Lorestan Province. Plant Pests Research, 4 (4), 25–34. [in Persian with English abstract]
Hashemi Khabir, Z., Haddad Irani Nejad, K. & Akbari, A. (2013) Some of edaphic mesostigmatic and prostigmatic mites in Southwest of East Azerbaijan Province, Iran. In: Joharchi, O. & Saboori, A. (Eds.), Program & Abstract Book of the Second International Persian Congress of Acarology. Faculty of Agriculture, University of Tehran, Karaj, p. 13.
Hirschmann W. & Krauss, W. (1965) Gangsystematik der Parasitiformes. Teil 8. Gamasiden. Bestimmungstafeln von 55 Pachylaelaps-Arten. Acarologie, Schriftenreihe für vergleichende Milbenkunde, 7, 1–5, pls. 1–28.
Huhta, V. (2016) Catalogue of the Mesostigmata mites in Finland. Memoranda: Societatis pro Fauna et Flora Fennica, 92, 129–148.
Huhta, V., Siira-Pietikäinen, A., Penttinen, R. & Räty, M. (2010) Soil fauna of Finland: Acarina, Collembola and Enchytraeidae. Memoranda: Societatis pro Fauna et Flora Fennica, 86, 59–82.
Hyatt, K.H. (1956) I.— British mites of the genus Pachyseius Berlese, 1910 (Gamasina-Neoparasitidae). Annals and Magazine of Natural History, Series 12, 9 (97), 1–6.
https://doi.org/10.1080/00222935608655716
Jalaeian, M., Saboori, A. & Seyedoleslami, H. (2004) Report of mesostigmatic mites in Iran. Journal of the Entomological Society of Iran, 23 (2), 125–126.
Kamali, K., Ostovan, H. & Atamehr, A. (2001) A Catalog of Mites & Ticks (Acari) of Iran. Islamic Azad University Scientific Publication Center, Tehran, 192 pp.
Karg, W. (1971) Acari (Acarina), Milben, Unterordnung Anactinochaeta (Parasitiformes). Die freilebenden Gamasina (Gamasides), Raubmilben. Die Tierwelt Deutschlands, 59, 1–475.
Karg, W. (1993) Acari (Acarina), Milben. Parasitiformes (Anactinochaeta). Cohors Gamasina Leach. Raubmilben. 2. Überarbeitete Auflage. Die Tierwelt Deutschlands, 59, 1–523.
Kazemi, Sh. & Rajaei, A. (2013) An annotated checklist of Iranian Mesostigmata (Acari), excluding the family Phytoseiidae. Persian Journal of Acarology, 2 (1), 63‒158.
https://doi.org/10.22073/pja.v2i1.9950
Keum, E., Takaku, G., Lee, K. & Jung, C. (2016) New records of phoretic mites (Acari: Mesostigmata) associated with dung beetles (Coleoptera: Scarabaeidae) in Korea and their ecological implication. Journal of Asia-Pacific Entomology, 19 (2), 353–357.
https://doi.org/10.1016/j.aspen.2016.04.002
Khademi, N., Saboori, A. & Faraji, F. (2006) Fauna of Mesostigmata in citrus orchards in Jahrom region, Iran. In: Bruin, J. (Ed.), Abstract Book of 12th International Congress of Acarology. Amsterdam, p. 91.
Khanjani, M. & Kamali, K. (2000a) A study of Acari fauna of clover (Trifolium sp.) in Hamadan. In: Proceeding of the 14th Iranian Plant Protection Congress. Vol. 1. Pests. Isfahan University of Technology, Isfahan, p. 234.
Khanjani, M. & Kamali, K. (2000b) Mites (Acari) associated with beans (Phaseolus volgaris L.) in Hamadan province. In: Proceeding of the 14th Iranian Plant Protection Congress, Vol. 1. Pests. Isfahan University of Technology, Isfahan, p. 246.
Khorsand, A., Ostovan, H., Amin, Gh.A. & Khorsand, Z. (2012) Study on fauna of mesostigmata mites (Acari: Mesostigmata) associated with cotton fields in Darab region. In: Sarafrazi, A., Asef, M.R., Mozhdehi, M., Mozhdehi, M., Solhjouy Fard, S. & Abdollahi, T. (Eds.), Proceedings of the 20th Iranian Plant Protection Congress. Shiraz University, Shiraz, p. 513.
Koroleva, E.V. (1976) New species of mites of the genus Olopachys Berlese, 1910 (Parasitiformes, Pachylaelaptidae). Entomologicheskoe Obozrenie, 55, 480–496.
Koroleva, E.V. (1977a) Novye vidy kleshchei roda Pachylaelaps Berlese, 1888 (Parasitiformes, Pachylaelaptidae). Parazitologicheskii Sbornik, 27, 119–148.
Koroleva, E.V. (1977b) Family Pachylaelaptidae Vitzthum, 1931. In: Ghilyarov M.S. & Bregetova. N.G. (Eds.), A key to the soil-inhabiting mites, Mesostigmata. Zoological Institute of the Academy of Sciences, Petrograd and USSR, Nauka, Leningrad, pp. 411‒483.
Lindquist, E.E. & Evans, G.O. (1965) Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata). Memoirs of the Entomological Society of Canada, 97 (S47), 5–64.
https://doi.org/10.4039/entm9747fv
Mahpikaran, M., Kazemi, Sh. & Bahrami, F. (2012) Edaphic mesostigmatic mites (Acari: Mesostigmata) of Bojnourd region. In: Sarafrazi, A., Asef, M.R., Mozhdehi, M., Mozhdehi, M., Solhjouy Fard, S. & Abdollahi, T. (Eds.), Proceedings of the 20th Iranian Plant Protection Congress. Shiraz University, Shiraz, p. 473.
Maleki, Sh., Ostovan, H., Baniameri, V. & Joharchi, O. (2016) Biodiversity of mesostigmatic soil mite fauna (Acari: Mesostigmata) of a city park located in Tehran, Iran. Journal of Entomological Society of Iran, 36 (3), 181−194. [in Persian with English abstract]
Mašán, P. (2007) A Review of the Family Pachylaelapidae in Slovakia, with Systematics and Ecology of European Species (Acari: Mesostigmata: Eviphidoidea). Institute of Zoology, Slovak Academy of Sciences, Bratislava, 247 pp.
Mašán, P. (2014) A review of the genus Pseudolaelaps Berlese, 1916 (Acari: Mesostigmata, Pseudolaelapidae), with descriptions of eleven new species from Europe. Systematic & Applied Acarology, 19 (3), 283–312.
https://doi.org/10.11158/saa.19.3.4
Mašán, P. (2018) A morphological re-evaluation of Pachyseius humeralis Berlese, 1910 (Acari, Mesostigmata, Pachylaelapidae). ZooKeys, 790, 35–44.
https://doi.org/10.3897/zookeys.790.26894
Mašán, P. & Halliday, B. (2009) Mesostigmatid mites associated with the dung beetle Copris lunaris (Coleoptera: Scarabaeidae). European Journal of Entomology, 106 (4), 545–550.
https://doi.org/10.14411/eje.2009.068
Mašán, P. & Halliday, B. (2014) Review of the mite family Pachylaelapidae (Acari: Mesostigmata). Zootaxa, 3776 (1), 1–66.
https://doi.org/10.11646/zootaxa.3776.1.1
Mašán, P. & Mihál, I. (Eds.) (2009) Arachnids of the Cerová vrchovina highland (Arachnida: Araneae, Pesudoscorpiones, Opiliones, Acari). Štátna ochrana prírody SR, Banská Bystrica- Správa CHKO Cerová vrchovina, Rimavská Sobota- Ústav zoológie SAV, Bratislava- Ústav ekológie lesa SAV, Zvolen, 311 pp.
Mašán, P., Mojahed, S., Hajizadeh, J., Hosseini, R. & Ahadiyat, A. (2018) On remarkable Pachylaelaps species with unusual sperm induction system in females (Acari, Mesostigmata, Pachylaelapidae). Systematic and Applied Acarology, 23 (9), 1726–1740.
https://doi.org/10.11158/saa.23.9.2
Masnavipour, M., Kazemi, Sh., Latifi, M. & Ziaaddini, M. (2014) Fauna of edaphic mites of the subcohort Dermanyssiae (Mesostigmata: Gamasina) in northwestern area of Kerman Province. Taxonomy and Biosystematics, 6 (20), 37–48. [in Persian with English abstract]
Moghimi, F., Ahadiyat, A., Kiadaliri, H. & Karaca, M. (2019) A new species of the genus Olopachys (Mesostigmata: Pachylaelapidae) from Mazandaran Province, Iran, with a review of the Iranian species. International Journal of Acarology, 45 (8), 428–437.
https://doi.org/10.1080/01647954.2019.1668477
Mohammad-Dustar-Sharaf, M., Shirdel, D. & Mirfakhraie, Sh. (2016) Introduction to some edaphic mesostigmatic mites (Acari: Mesostigmata) from Arasbaran forests, north of East Azerbaijan Province. Journal of Applied Researches in Plant Protection, 5 (2), 227–242. [in Persian with English abstract]
Mojahed, S., Hajizadeh, J., Hosseini, R. & Ahadiyat, A. (2017) Contribution to the Pachylaelapidae Berlese fauna (Acari: Mesostigmata) in some parts of Guilan province. Persian Journal of Acarology, 6 (4), 269–285.
https://doi.org/10.22073/pja.v6i4.29712
Mojahed, S., Hajizadeh, J., Hosseini, R. & Ahadiyat, A. (2019) A new species of Olopachys Berlese (Acari: Pachylaelapidae) from Iran with a key to the world species. Acarologia, 59 (1), 46–56.
https://doi.org/10.24349/acarologia/20194313
Moraza, M.L. & Johnston, D.E. (1990) Pachyseiulus hispanicus n. gen., n. sp., from Navarra (Northern Spain) (Acari: Mesostigmata: Pachylaelapidae). International Journal of Acarology, 16 (4), 213–218.
https://doi.org/10.1080/01647959008683870
Moraza, M.L. & Peña, M.A. (2005) The family Pachylaelapidae Vitzthum, 1931 on Tenerife Island (Canary Islands), with description of seven new species of the genus Pachylaelaps (Acari, Mesostigmata: Pachylaelapidae). Acarologia, 45 (2‒3), 103–129.
Nazari-Tajani, M., Hajizadeh, J. & Noei, J. (2012) First report of one pachylaelapid species (Mesostigmata: Pachylaelapidae) from Iran. In: Sarafrazi, A., Asef, M.R., Mozhdehi, M., Mozhdehi, M., Solhjouy Fard, S. & Abdollahi, T. (Eds.), Proceedings of the 20th Iranian Plant Protection Congress. Shiraz University, Shiraz, p. 510.
Nemati, A., Riahi, E., Khalili-Moghadam, A. & Gwiazdowicz, D.J. (2018) A catalogue of the Iranian Mesostigmata (Acari): additions and updates of the previous catalogue. Persian Journal of Acarology, 7 (2), 115–191.
https://doi.org/10.22073/pja.v7i2.36985
Oudemans, A.C. (1902) Verslag. Tijdschrift voor Entomologie, 45, 50–64.
Özbek, H.H. (2014) A new species of Olopachys Berlese (Acari: Pachylaelapidae) from northeastern Turkey. International Journal of Acarology, 40 (4), 328–331.
https://doi.org/10.1080/01647954.2014.916748
Rahmani, H., Kamali, K., Saboori, A. & Nowzari, J. (2006) Mites of the honeybee (Apis mellifera L.) hives in Tehran province (Iran). In: Manzari, S. (Ed.), Proceedings of the 17th Iranian Plant Protection Congress. Vol. 1. Pests. University of Tehran, Karaj, p. 195.
Rajaei, A., Kazemi, Sh. & Yazdanian, M. (2011) Soil-inhabiting Mesostigmatia of the superfamilies Uropodoidea and Eviphidoidea (Acari) in Baft region, Kerman province, Iran. In: Kazemi, Sh. & Saboori, A. (Eds.), Abstract and Proceeding Book of the First Persian Congress of Acarology. International Center for Science, High Technology & Environmental Sciences. Kerman, p. 50.
Rostami, E., Abassipour, H., Khanjani, M. & Askarianzadeh, A. (2010) Faunistic study of fruit tree mites in Hamedan Province, west of Iran. In: de Moraes, G.J., Castilho, R.C. & Flechtmann, C.H.W. (Eds.), Abstract Book of the 13th International Congress of Acarology. PE, Recife, p. 233.
Şahin, G. & Özbek, H.H. (2018) Mites of the family Pachylaelapidae in Bayburt Province, Turkey (Acari: Mesostigmata), with a new record and three new species. Zootaxa, 4514 (2), 243–255.
https://doi.org/10.11646/zootaxa.4514.2.7
Salarzehi, S., Hajiqanbar, H., Faraji, F., Joharchi, O., Noei, J. & Oliyai, A. (2011) Fauna of some mesostigmatic mites (Acari: Mesostigmata) in Kashmar region, Iran. In: Kazemi, Sh. & Saboori, A. (Eds.), Abstract and Proceeding Book of the First Persian Congress of Acarology. International Center for Science, High Technology & Environmental Sciences, Kerman, p. 56.
Salmane, I. (2001) A check-list of Latvian Gamasina mites (Acari, Mesostigmata) with short notes to their ecology. Latvijas Entomologs, 38, 27–38.
Salmane, I. & Brumelis, G. (2010) Species list and habitat preference of Mesostigmata mites (Acari, Parasitiformes) in Latvia. Acarologia, 50 (3), 373–394.
https://doi.org/10.1051/acarologia/20101978
Salmane, I. & Kontschán, J. (2005) Soil Gamasina mites (Acari, Parasitiformes, Mesostigmata) from Hungary. I. Latvijas Entomologs, 42, 48–56.
Schrader, S. & Bayer, B. (2000) Abundances of mites (Gamasina and Oribatida) and biotic activity in arable soil affected by tillage and wheeling. Braunschweiger Naturkundliche Schriften, 6 (1), 165–181.
Schweizer, J. (1961) Die Landmilben der Schweiz (Mittelland, Jura und Alpen). Parasitiformes Reuter. Denkschriften der Schweizerischen Naturforschenden Gesellschaft, 84, 1–207.
Seifori, M., Saboori, A. & Vafai Shooshtari, R. (2006) A faunistic survey of eviphidoid mites (Mesostigmata: Eviphidoidea) in the soil of Arak region, Markazi province, Iran. In: Bruin, J. (Ed.), Abstract Book of 12th International Congress of Acarology. Amsterdam, p. 187.
Sellnick, M. (1969) Eine neue Pachylaelaps-Art von Island. Entomologisk Tidskrift, 90 (1–2), 131–133.
Shariati, R., Jalalizand, A. & Moradi-Faradonbeh, M. (2017) The fauna of edaphic mites (Acari: Mesostigmata) in Saman County, Southwest Iran. In: Hajiqanbar, H. & Saboori, A. (Eds.), Program & Abstract Book of the Third International Persian Congress of Acarology. University of Tehran, Tehran, p. 36.
Soleimani, M., Ostovan, H. & Joharchi, O. (2011) Mesostigmatic mites (Acari: Mesostigmata) in Marvdasht region, Fars province, Iran. In: Kazemi, Sh. & Saboori, A. (Eds.), Abstract and Proceeding Book of the First Persian Congress of Acarology. International Center for Science, High Technology & Environmental Sciences. Kerman, p. 17.
Valizadeh, S., Ahadiyat, A., Bagheri, M. & Joharchi, O. (2019) Second world record of Olopachys hallidayi Özbek from Iran (Mesostigmata: Pachylaelapidae). Persian Journal of Acarology, 8 (2), 169–174.
http://dx.doi.org/10.22073/pja.v8i2.43928
Willmann, C. (1940) Neue Milben aus Höhlen der Balkanhalbinsel, gesammelt von Prof. Dr. K. Absolon, Brünn (2. Mitteilung.). Zoologischer Anzeiger, 130 (9/10), 209–218.
Willmann, C. (1953) Neue Milben aus den östlichen Alpen. Sitzungsberichten der Österreichischen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse, I, 162 (6), 449–519.
Zakeri, V., Kamali, K. & Hajiqanbar, H.R. (2012) Coprophage and edaphic mites of the families Macrochelidae and Pachylaelapidae in eastern region of Golestan Province, Iran. Journal of Iranian Plant Pests Research, 1 (1), 17–23. [in Persian with English abstract]
Zarei, E. & Kazemi, Sh. (2014) Fauna of edaphic Mesostigmata (Acari) in apple orchards of Miandoab County, West Azerbaijan Province, Iran. In: 21st Iranian Plant Protection Congress. Urmia University, Urmia, p. 980.