Skip to main content Skip to main navigation menu Skip to site footer
Type: Monograph
Published: 2020-03-19
Page range: 1–78
Abstract views: 106
PDF downloaded: 1

The millipede family Conotylidae in northwestern North America, with a complete bibliography of the family (Diplopoda, Chordeumatida, Heterochordeumatidea, Conotyloidea)

Professor Emeritus, Hampden-Sydney College, Hampden-Sydney, VA 23943, present address: 1950 Price Drive, Farmville, VA 23901.
Santa Barbara Botanic Garden, 1212 Mission Canyon Road Santa Barbara, CA 93105-2126. Email address:
Department of Entomology, Texas A&M University, TAMU 2475, College Station, TX 77843.
Arthropoda new genera new species descriptions California Nevada Oregon Idaho Washington British Columbia Alberta

Abstract

The center of diversity for the chordeumatidan millipede family Conotylidae is northwestern North America, where five of six subfamilies and ten of fourteen previously described genera occur; in this paper, seven additional new genera and 33 new species from the region are described. New species in the genus Taiyutyla Chamberlin, 1952 are: Taiyutyla tillamook, Taiyutyla acuphora, and Taiyutyla amicitia; in the genus Bollmanella Chamberlin, 1941: Bollmanella bombus, Bollmanella washingtonensis and Bollmanella leonardi; in the genus Brunsonia Loomis & Schmitt, 1971: Brunsonia pulchra, Brunsonia digitata, Brunsonia wenatchee, Brunsonia chelanoparva, Brunsonia chelanomagna, Brunsonia selwayana and Brunsonia benewah. Vancouvereuma n. gen. is described with the type species Taiyutyla shawi Shear 2004. Calityla n. gen. includes the new species Calityla siskiyou, Calityla ubicki, Calityla trinitaria, Calityla essigi, and Calityla humboldtensis. Ovaskella, n. gen. includes the new species Ovaskella ovaskae and Ovaskella sinuosa. Karagama, n. gen. includes the new species Karagama ladybird. Complicatella, n. gen. includes the new species Complicatella pectenifera and Complicatella neili. Bifurcatella n. gen. includes the new species Bifurcatella olympiana, Bifurcatella hoh, Bifurcatella angulata, Bifurcatella pacifica, Bifurcatella germania, Bifurcatella uniclada, Bifurcatella inflata and Bifurcatella hobo. Loomisiella, n. gen. includes the new species Loomisiella evergreen and Loomisiella pylei. New distribution records are provided for numerous previously described species of conotylids. A complete bibliography of the family Conotylidae of the world is also included.

 

References

  1. Bollman, C.H. (1889) Catalog of the myriapods of Indiana. Proceedings of the United States National Museum, 11, 403–410.

    https://doi.org/10.5479/si.00963801.11-731.403

    Bollman, C.H. (1893) The Myriapoda of the United States. United States National Museum Bulletin, 43, 1–210.

    Buckett, J.S. & Gardner, M.R. (1967) A new family of cavernicolous millipedes with the description of a new genus and species from Idaho (Diplopoda: Chordeumida: Chordeumidea). The Michigan Entomologist, 1, 117–126.

    Causey, N.B. (1961) Austrotyla, a new millipede genus (Chordeumida: Conotylidae: Conotylinae). Proceedings of the Biological Society of Washington, 74, 251–266.

    Causey, N.B. (1972) Two new conotylid millipeds from western North America and a key to the genus Adrityla (Chordeumida: Diplopoda). Proceedings of the Louisiana Academy of Sciences, 35, 27–32.

    Chamberlin, R.V. (1910) Diplopoda from the western states. Annals of the Entomological Society of America, 3, 233–262.

    https://doi.org/10.1093/aesa/3.4.233

    Chamberlin, R.V. (1920) Some records of Canadian myriopods. The Canadian Entomologist, 52 (4), 94–95.

    Chamberlin, R.V. (1941) New western millipeds. Bulletin of the University of Utah, Biological Series, 6, 3–23.

    Chamberlin, R.V. (1951) Eleven new western millipeds. Natural History Miscellanea of the Chicago Academy of Sciences, 87, 1–12.

    Chamberlin, R.V. (1952) Two Oregon millipeds of the order Chordeumida. Natural History Miscellanea of the Chicago Academy of Sciences, 113, 1–3.

    Cook, O.F. (1896) On recent diplopod names. Brandtia, 2, 5–8.

    Cook, O.F. & Collins, G. (1895) The Craspedosomatidae of North America. Annals of the New York Academy of Sciences, 9, 1–100.

    https://doi.org/10.1111/j.1749-6632.1896.tb55430.x

    Golovatch, S.I. (1980) New forms of Diplopoda from the Soviet Far East and their zoogegraphical relationships. Zoologicheskii zhurnal, 59, 199–207.

    Golovatch, S.l. & Mikhaljova, E.Y. (1983) A new family of East-Palaearctic Chordeumida (Diplopoda), with a description of a new genus and a new species. Bulletin of the Moscow Society of Naturalists, Biological Series, 83 (4), 66–71. [in Russian]

    Harger, O. (1872) Descriptions of new North American myriapods. American Journal of the Sciences and Arts, Series III, 4, 117–121.

    https://doi.org/10.2475/ajs.s3-4.20.117

    Hoffman, R.L. (1961) Systematic and morphological notes on North American conotyloid diplopods. Transactions of the American Entomological Society, 87, 259–272.

    Hoffman, R.L. (1979) Classification of the Diplopoda. Muséum naturelle Genève, Genève, 237 pp.

    Hoffman, R.L. (1999) Checklist of the Millipeds of North and Middle America. Virginia Museum of Natural History Special Publication 8. Virginia Museum of Natural History, Martinsburg, 584 pp.

    Koch, C.L. (1847) System der Myriapoden. Herrich-Schäffer, Kritische Revision der Insectfauna Deutschlands, 3, 1496.

    Loomis, H.F. (1939) The millipeds collected in Appalachian caves by Mr. Kenneth Dearolf. Bulletin of the Museum of Comparative Zoology, 86, 165–193.

    Loomis, H.F. (1943) New cave and epigean millipeds. Bulletin of the Museum of Comparative Zoology, 92, 373–410.

    Loomis, H.F. & Schmitt, R. (1971) The ecology, distribution and taxonomy of the millipeds of Montana west of the continental divide. Northwest Science, 45, 107–131.

    Mauries, J.-P. (1978) Le genre neozelandais Schedotrigona Silvestri, 1903: revision et place dans une nouvelle classification des Craspedosomides (Myriapoda, Diplopoda, Craspedosomida). Bulletin Muséum national de histoire naturelle, Series 3, 510, 43–66.

    Mauries, J.-P. (1982) Une espèce nouvelle du genre Lankasoma (Diplopoda, Craspedosomida, Lankasomidae), avec notes sur quleques Chordeumida recement decrits. Steenstrupia, 8, 177–180.

    McNeill, J. (1887) Descriptions of twelve new species of Myriapoda, chiefly from Indiana. Proceedings of the United States National Museum, 10, 328–334.

    https://doi.org/10.5479/si.00963801.10-632.328

    Morrison, R.B. (1963) Lake Lahontan: geology of southern Carson Desert, Nevada. Geological Survey Professional Paper, 401, 1–156.

    https://doi.org/10.3133/pp401

    Richart, C.H. & Hedin, M. (2013) Three new species in the harvestmen genus Acuclavella (Opiliones, Dyspnoi, Ischyropsalidoidea), including description of male Acuclavella quattuor Shear, 1986. ZooKeys, 311, 19–68.

    https://doi.org/10.3897/zookeys.311.2920

    Shear, W.A. (1971) The millipede family Conotylidae in North America, with a description of the new family Adritylidae (Diplopoda: Chordeumida). Bulletin of the Museum of Comparative Zoology, 141, 55–96.

    Shear, W.A. (1972) Studies in the millipede order Chordeumida (Diplopoda): a revision of the family Cleidogonidae and a reclassification of the order in the New World. Bulletin of the Museum of Comparative Zoology, 144, 151–352.

    Shear, W.A. (1974) The milliped genus Bollmanella (Diplopoda, Chordeumida, Conotylidae). Psyche, 81, 134–146.

    https://doi.org/10.1155/1974/51907

    Shear, W.A. (1976) The milliped family Conotylidae (Diplopoda, Chordeumida): revision of the genus Taiyutyla, with notes on recently proposed taxa. American Museum Novitates, 2600, 1–22.

    Shear, W.A. (1987) Systematic position of the millipede Japanosoma scabrum Verhoeff (Chordeumatida, Conotylidae). Myriapodologica, 2, 21–27.

    Shear, W.A. (2000) On the millipede family Heterochordeumatidae, with comments on the higher classification of the order Chordeumida. Invertebrate Taxonomy, 14, 363–376.

    https://doi.org/10.1071/IT99016

    Shear, W.A. (2004) Two new species in the millipede genus Taiyutyla from British Columbia, Canada, and Washington, USA. Myriapodologica, 8, 13–20.

    Shear, W.A. (2007) Cave millipeds of the United States. V. The genus Idagona Buckett & Gardner (Chordeumatida, Conotylidae, Idagoninae). Zootaxa, 1463 (1), 1–12.

    https://doi.org/10.11646/zootaxa.1463.1.1

    Shear, W.A. & Krejca, J.K. (2011) Cave millipeds of the United States. IX. A new species of the genus Taiyutyla (Diplopoda, Chordeumatida, Conotylidae) from caves in Sequoia and Yosemite National Parks, California, USA. Journal of Cave and Karst Studies, 73, 93–98.

    https://doi.org/10.4311/jcks2010lsc157

    Shear, W.A. & Shelley, R.M. (2007) Tingupa tlingitorum, n. sp., a new millipede from Haines, Alaska, USA, with notes on the generic distribution and a revised key to species (Chordeumatida, Tingupidae). Zootaxa, 1393 (1), 53–59.

    https://doi.org/10.11646/zootaxa.1393.1.6

    Shear, W.A. & Steinmann, D.B. (2013) Cave millipedes of the United States. XIII. A new, troglobiotic species of Austrotyla from Colorado (Diplopoda, Chordeumatida, Conotylidae). Zootaxa, 3745, 486–490.

    Shear, W.A. & Tsurusaki, N. (1995) Japanese chordeumatid millipeds. III. Yasudatyla, a new genus of alpine conotylid millipedes (Diplopoda, Chordeumatida, Conotylidae). Myriapodologica, 3, 98–106.

    Shelley, R.M. (1993) The myriapod types of Oscar Harger (Arthropoda: Diplopoda, Chilopoda). Brimleyana, 18, 1–13.

    Shelley, R.M. (1997) Corypus kavanaughi, n. sp., a large-bodied chordeumatoid milliped from Idaho, with a review of the genus (Conotylidae). Myriapodologica, 4, 121–126.

    Shelley, R.M. (2002) The millipeds of central Canada (Arthropoda: Diplopoda), with reviews of the Canadian fauna and diplopod faunistic studies. Canadian Journal of Zoology, 80, 1863–1875.

    https://doi.org/10.1139/z02-170

    Shelley, R.M., Medrano, M.F. & Ovaska, K. (2009) The millipede family Tingupidae (Chordeumatida) on Kodiak Island, Alaska, USA, a geographically remote record of indigenous Diplopoda. Insecta Mundi, 105, 1–5.

    Shelley, R.M., Medrano, M.F., Shear, W.A., Ovaska, K., White, K.J. & Havard, E.I. (2009) Distribution extensions of the millipede families Conotylidae and Rhiscosomididae (Diplopoda: Chordeumatida) into northern British Columbia and southern Alaska. Insecta Mundi, 71, 1–6.

    Shelley, R.M., Shear, W.A., Leonard, W.P. & Ovaska, K. (2007) Diplopoda, Chordeumatida, Caseyidae, Opiona columbiana Chamberlin, 1951: Distribution extensions into the Alexander Archipelago, Alaska, USA, Queen Charlotte Islands, British Columbia, Canada, and eastern & southern Washington State, USA; additional new records from British Columbia and Washington. Check List, 3, 13–17.

    https://doi.org/10.15560/3.1.14

    Verhoeff, K.-W. (1914) Ascospermopheren aus Japan. Zoologischer Anzeiger, 43, 342–370.

    Verhoeff, K.-W. (1932) Diplopoden-beitrage (124. Diplopoden aufsatz). Zoologisches Jahrbücher, Systematisches Arbeiten, 62, 469–524, pls. 1–2.

    Wynne, J.J. & Shear, W.A. (2016) A new millipede, Austrotyla awishishola, n. sp. (Diplopoda, Chordeumatida, Conotylidae) from New Mexico, USA, and the importance of cave moss gardens as refugial habitats. Zootaxa, 4084 (2), 285–292.

    https://doi.org/10.11646/zootaxa.4084.2.8