Abstract
Japanese species and genera of the tribe Smicromyrmini Bischoff, 1920 are revised. Most of the relevant types were examined. The following eight species in the genera Smicromyrme Thomson, 1870, Nemka Lelej, 1985, Mickelomyrme Lelej, 1995, Andreimyrme Lelej, 1995, Ephucilla Lelej, 1995 and Sinotilla Lelej, 1995 are recognized from Japan: Sm. lewisi Mickel, 1935, ♂♀; N. yasumatsui (Mickel, 1936), stat. rev. et comb. nov., ♂♀; M. hageni (Zavattari, 1913), ♂♀; A. substriolata (Chen, 1957), ♀ (Amami-ôshima Is., Okinawa-jima Is.); E. yoshitakei sp. nov., ♂♀ (Amami-ôshima Is.), E. suginoi sp. nov., ♂♀ (Okinawa-jima Is.), E. brevitegula sp. nov., ♂♀ (Ishigaki-jima Is., Iriomote-jima Is.); Si. jambar sp. nov., ♂ (Okinawa-jima Is.). The genera Andreimyrme and Sinotilla are newly recorded from Japan. Nemka wotani (Zavattari, 1913) and Ephucilla naja (Zavattari, 1913) are excluded from the Japanese fauna. New synonymies are proposed for N. wotani (Zavattari, 1913) [=Smicromyrme chihpenchia Tsuneki, 1993, syn. nov.], N. limi limi (Chen, 1957) [=Smicromyrme limi nanhai Chen, 1957, syn. nov.], A. substriolata [=Smicromyrme tridentiens Chen, 1957, syn. nov.] and Neotrogaspidia pustulata (Smith, 1873) [=Smicromyrme yakushimensis Yasumatsu, 1934, syn. nov.]. Phimenes flavopictus formosanus (Zimmermann, 1931) (Hymenoptera: Vespidae: Eumeninae) is recorded as a host of Ephucilla brevitegula. Other new records include: Mickelomyrme athalia (Pagden, 1934) from China (Guangdong, Fujian, Yunnan) and Vietnam (Hoa Binh), M. bakeri (Mickel, 1934) from Malaysia (Borneo) and Indonesia (Sulawesi), M. pusillaeformis (Hammer, 1962) from Indonesia (Bali), Andreimyrme sarawakensis Lelej, 1996 from Peninsular Malaysia (Kuala Lumpur) and A. substriolata from Laos (Xiengkhouang, Houapanh), Thailand (Chiang Mai), Peninsular Malaysia (Kuala Terengganu) and Indonesia (Sumatra). Distributions of Japanese smicromyrmine species are classified into the following three patterns: Asian continental-Palaearctic Japan (Sm. lewisi and N. yasumatsui), Taiwan-Ryûkyû (M. hageni, E. yoshitakei, E. suginoi, E. brevitegula, Si. jambar) and Asian continental-Ryûkyû (A. substriolata). Species of Asian continental-Palaearctic Japan are widely distributed in Palaearctic part of Japan and eastern part of Eurasia, or have the closest relative in continental Asia. They are considered to be native to Japan since it was a part of Eurasia, or have immigrated to Japan through the land bridge between Korean Peninsula and Japan by the end of the Upper Pleistocene. The Taiwan-Ryûkyû species are immigrants from Taiwan through the land bridge. The Asian continental-Ryûkyû species is presumed to have immigrated to Japan during the Upper Pleistocene when the eastern edge of Eurasian Continent was close to Amami-ôshima Is.
References
Aguiar, A.P., Deans, A.R., Engel, M.S., Forshage, M., Huber, J.T., Jennings, J.T., Johnson, N.E., Lelej, A.S., Longino, J.T., Lohrmann, V., Mikó, I., Ohl, M., Rasmussen, C., Taeger, A. & Yu, D.S.K. (2013) Order Hymenoptera. In: Zhang, Z.-Q. (Ed.), Animal Biodiversity: An Outline of Higher-level Classification and Survey of Taxonomic Richness (Addenda 2013). Zootaxa, 3703 (1), 51–62.
https://doi.org/10.11646/zootaxa.3703.1.12
Amini, A., Lelej, A.S., Sadegh, H. & Karimi, J. (2014) First record of the velvet ants (Hymenoptera: Mutillidae) reared from puparia of the ber fruit fly Carpomya vesuviana Costa (Diptera: Tephritidae) in Iran. Zootaxa, 3861 (6), 585–590.
https://doi.org/10.11646/zootaxa.3861.6.5
André, E. (1894) Contribution a la connaissance des Mutilles de l’Inde. Journal of the Bombay Natural History Society, 8, 462–484.
André, E. (1896) Mutillides nouveaux ou imparfaitement connues fouisant pourtie des collections du muscc National de Hongrie. Termeszetraizi fuzetek, 19 (1), 9–25.
André, E. (1898) Étude sur les mutillides du Muséum de Paris. Annales de la Société Entomologique de France, 67, 1–79.
André, E. (1899–1903) Species des Hyménoptères d’Europe & d’Algérie. Tome Huitième. Les Mutillides. Vve Dubosclard and A. Hermann, Paris, 480 pp., 15 pls. [pp. 1–64 + pls. 1–8 (1899), pp. 65–144 (1900), pp. 145–304 (1901), pp. 305–384 (1902a), 385–478 (1903a), pls. 9–15 (year unknown)]
André, E. (1902b [1903b]) Family Mutillidae. In: Wytsman, P. (Ed.), Genera Insectorum. Fascicles 11. L. Desmet-Verteneuil, Bruxelles, pp. 1–77, 3 pls.
André, E. (1905) Mutillides nouveaux ou peu connues de l’Ancien Monds et de l’Australie. Zeitschrift für systematische Hymenopterologie und Dipterologie, 5 (1–6), 201–216 + 265–277.
Ashmead, W.H. (1899) Superfamilies in the Hymenoptera and generic synopses of the families Thynnidae, Myrmosidae, and Mutillidae. Journal of the New York Entomological Society, 7, 45–60.
Ashmead, W.H. (1904) Descriptions of new genera and species of Hymenoptera from the Philippine Islands. Proceedings of the United States National Museum, 28 (1387), 127–158.
https://doi.org/10.5479/si.00963801.28-1387.127
Azuma, S., Yafuso, M., Kinjo, M., Hayashi, M., Kohama, T., Sasaki, T., Kimura, M. & Kawamura, F. (2002) Check list of the insect of the Ryukyu Islands (Second Edition). The Biological Society of Okinawa, Okinawa, 570 pp. [in Japanese]
Barthélémy, C., Lelej, A.S. & Loktionov, V.M. (2018) A contribution to the Mutillidae (Hymenoptera) of Hong Kong. Zootaxa, 4514 (2), 230–242.
https://doi.org/10.11646/zootaxa.4514.2.6
Bartholomay, P.R., Williams, K.A., Luz, D.R. & Morato, E.F. (2015) Frigitilla gen. nov., a new genus of Amazonian Mutillidae (Hymenoptera). Zootaxa, 3957 (1), 49–58.
https://doi.org/10.11646/zootaxa.3957.1.3
Bayliss, P.S. & Brothers, D.J. (2001) Behaviour and host relationships of Dolichomutilla sycorax (Smith, 1855) (Hymenoptera: Mutillidae, Sphecidae). Journal of Hymenoptera Research, 10, 1–9.
Bischoff, H. (1920–1921) Monographie der Mutilliden Afrikas. Archiv für Naturgeschichte, 86A (1–3) & 86A (4–5), 1–480 (1920) & 481–830 (1921).
Bogusch, P. (2006) The velvet ants (Hymenoptera: Mutillidae) of the Czech Republic and Slovakia: an identification key and annotated checklist. Acta Musei Moraviae, Scientiae biologicae, Brno, 91, 103–148.
Brothers, D.J. (1975) Phylogeny and classification of the aculeate Hymenoptera, with special reference to Mutillidae. The University of Kansas Science Bulletin, 50 (11), 483–648.
Brothers, D.J. (1989) Alternative life-history styles of mutillid wasps (Insecta, Hymenoptera). In: Bruton, M.N. (Ed.), Alternative Life-History Styles of Animals. Perspectives in Vertebrate Sciences 6. Springer, Dordrecht, pp. 279–291.
https://doi.org/10.1007/978-94-009-2605-9_14
Brothers, D.J. (1995) Mutillidae. In: Hanson, P.E. & Gauld, I.D. (Ed.), The Hymenoptera of Costa Rica. Oxford University Press, Oxford, pp. 541–548.
Brothers, D.J. (2018) Aglaotilla, a new genus of Australian Mutillidae (Hymenoptera) with metallic coloration. Zootaxa, 4415 (2), 357–368.
https://doi.org/10.11646/zootaxa.4415.2.6
Brothers, D.J. & Lelej, A.S. (2017) Phylogeny and higher classification of Mutillidae (Hymenoptera) based on morphological reanalyses. Journal of Hymenoptera Research, 60, 1–97.
https://doi.org/10.3897/jhr.60.20091
Brothers, D. J., Lelej, A. S. & Williams, K. A. (2019) Genus-group names of Mutillidae (Hymenoptera): corrections and updates since 2008. Zootaxa, 4651 (3), 578–588.
https://doi.org/10.11646/zootaxa.4651.3.10
Brothers, D.J., Tschuch, G. & Burger, F. (2000) Associations of mutillid wasps (Hymenoptera, Mutillidae) with eusocial insects. Insectes Sociaux, 47, 201–211.
https://doi.org/10.1007/PL00001704
Cameron, P. (1892) Hymenoptera Orientalis, or contributions to a knowledge of the Hymenoptera of the Indian Zoological Region. Part IV. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 35, 97–137, 1 pl.
Cameron, P. (1898) Hymenoptera Orientalia, or Contributions to a knowledge of the Hymenoptera of the Oriental Zoological Region. Part VII. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 42 (11), 1–84, 4 pls.
Cameron, P. (1900) Hymenoptera Orientalia, or contributions to the knowledge of the Hymenoptera of the Oriental zoological region. Part IX. The Hymenoptera of the Khasia Hills. Part II. Section I. Memoirs and Proceedings of the Manchester Literary and Philosophical Society, 44 (15), 1–114.
Chen, C.-W. (1957) A revision of the velvety ants or Mutillidae of China (Hymenoptera). Quarterly Journal of the Taiwan Museum, 10 (3–4), 135–224, 6 pls.
Chu, Y.-I. & Yamanaka, T. (1973–1975) A check list of the present and old name of insect collected localities in Taiwan. Journal of the National Taiwan Museum, 16–18, 31–72 (1973), 51–75 (1974) & 121–150 (1975).
Dalla Torre, C.G. de (1897) Catalogus Hymenopterorum hucusque Descriptorum Systematicus et Synonimicus. Volumen VIII: Fossores (Sphegidae). Guilelmi Engelmann, Lipsiae, [4] + viii + 750 pp.
Dufour, L. (1838) Observations sur le genre Stizus. Annales de la Société Entomologique de France, 7, 269–279.
Dutt, G.R. (1912) Life histories of Indian insects IV (Hymenoptera). Memoirs of the Department of Agriculture in India, 12 (4), 183–267.
Fabricius, J.C. (1787) Mantissa insectorum sistens eorum species nuper detecta, adjectis characteribus genericis, differentiis specificis, emendationibus, observationibus. Tom. 1. Impensis Christ. Gottl. Proft, Hafniae (Copenhagen), 348 pp.
https://doi.org/10.5962/bhl.title.11657
Fabricius, J.C. (1793) Entomologica Systematica emendata et aucta, secundum Classes, Ordines, Genera, Species adjectis Synonymis, Locis, Observationibus, Descriptionibus. Tom. 2. Christ. Gottl. Proft, Hafniae (Copenhagen), viii + 519 pp.
https://doi.org/10.5962/bhl.title.122153
Fabricius, J.C. (1804) Systema Piezatorum secundum Ordines, Genera, Species adjectis Synonymis, Locis, Observationibus, Descriptionibus. Carolum Reichard, Brunsvigae, i-xiv, 15–440, 1–30.
https://doi.org/10.5962/bhl.title.10490
Frivaldszky, J. (1876) Data ad faunam Hungariae meridionalis comitatum Temes et Krassó. Mathematikai és Természetudomanyi Közlemények, 13, 285–378.
Fukasawa, S. & Miyano, S. (2010) Fauna of the aculeate Hymenoptera (Insecta) from the Izu Islands, Japan. Journal of the Natural History Museum and Institute, Chiba, 11 (1), 11–28. [in Japanese]
Gauld, I.D. & Mitchell, P.A. (1981) The taxonomy, distribution and host preferences of Indo-Papuan parasitic wasps of the subfamily Ophioninae (Hymenoptera: Ichneumonidae). CAB: Slough. Commonwealth Institute of Entomology, London, 611 pp.
Gôukon, K. (1993) Host records and some bionomical notes on mutillid wasps in Japan (Hymenoptera, Mutillidae). Chugoku Kontyu, 7, 35–40. [in Japanese]
Grandi, G. (1961) Studi di un Entomologo Sugli Imenotteri Superiori. Bolletino dell’Istituto di Entomologia della Università Degli Studi di Bologna, 25, i–xv + 1–659.
Gyotoku, N. (1941) Record of Smicromyrme lewisi yanoi Mickel collected at light. Mushi, 14 (1), 7. [in Japanese]
Hadley, A. (2008) CombineZM software. Available from: https://combinezm.en.lo4d.com/ (accessed 7 May 2019)
Hammer, K. (1960 [1962]) Mutilliden (Insecta: Hymenoptera) aus dem Indischen Museum in Calcutta. Records of the Indian Museum, 58 (1), 1–51.
Haneda, Y., Inoue, S., Nozaka, C., Tano, T., Kurokawa, H. & Murota, T. (2005) The Hymenoptera recorded at Mt. Monju, Fukui Prefecture, Central Japan. Tsunekibachi, 4, 53–75. [in Japanese]
Haneda, Y., Nozaka, C., Tano, T., Kurokawa, H. & Murota, T. (2006a) Aculeate Hymenoptera collected from Toyama Prefecture in 2005. Tsunekibachi, 8, 1–48. [in Japanese]
Haneda, Y., Nozaka, C., Tano, T., Kurokawa, H. & Murota, T. (2006b) The aculeate Hymenoptera collected at Tsushima, Nagasaki Prefecture in August of 2006. Tsunekibachi, 10, 39–52. [in Japanese]
Haneda, Y. & Yamamoto, E. (2000) Aculeata collected in Odamiyama and surrounding area (Hymenoptera). In: Yamamoto, E. (Ed.), Nature of Odamiyama. II. Oda-chô, Ehime, pp. 775–837. [in Japanese]
Hashimoto, Y. & Nakanishi, A. (1997) A list of the type-specimens of Hymenoptera described by K. Tsuneki in the Museum of Nature and Human Activities, Hyogo (MNHAH). Catalogue of Collections in the Museum of Nature and Human Activities, Hyogo No. 2. Museum of Nature and Human Activities, Sanda, 47 pp., 8 pls.
Hayashi, M., Kadowaki, H., Matsuda, T., Fukaya, H. & Chikami, Y. (2013) Research on insect distribution of Oki Islands, Shimane Prefecture, Japan. Part 3. Special Bulletin of the Hoshizaki Green Foundation, 8, 1–73.
Hikida, T. & Motokawa, J. (1999) Phylogeographical relationships of the skinks of the genus Eumeces (Reptilia: Scincidae) in East Asia. In: Ota, H. (Ed.), Tropical island herpetofauna: origin, current diversity, and conservation. Elsevier, Amsterdam, pp. 231–247.
Hisamatsu, M. (2004) List of Hymenoptera recorded in Ibaraki Prefecture. Bulletin of Ibaraki Nature Museum, 7, 125–164. [in Japanese]
Hymenoptera Anatomy Ontology (2013) Hymenoptera Glossary. Available from: http://glossary.hymao.org (accessed 7 May 2019)
Ikudome, S. & Yamane, Sk. (2009) Ants, wasps and bees of Take-shima, Northern Ryukyus, Japan (Hymenoptera, Aculeata). The Bulletin of the Institute of Minami-Kyûshû Regional Science, Kagoshima Women’s Junior College, 24, 1–8.
International Commission on Zoological Nomenclature (ICZN) (1999) International Code of Zoological Nomenclature. Fourth Edition. ITZN, London, xxx + 306 pp.
Invrea, F. (1940) Mutillidi dell’isola di Cipro raccolti dal Sig. G.A. Mauromoustakis. Bollettino della Società Entomologica Italiana, 72, 118–122.
Invrea, F. (1943) A proposito di una Smicromyrme del’Isola di Borneo nota tassonomica e combiamento di nome (Hymenoptera, Mutillidae). Annali del Museo Civico di Storia Naturale di Genova, 61, 191–193.
Invrea, F. (1951) Mutillidi nuovi o notevoli del Bacino Mediterraneo (Hymenoptera, Mutillidae). II Parte. Bolletino della Società Entomologica Italiana, 81 (3–4), 34–43.
Invrea, F. (1964) Mutillidae-Myrmosidae. Fauna d’Italia, 5. Edizioni Calderini, Bologna, xii + 302 pp. + [1].
Ishikawa, R. (1965) Family Mutillidae. In: Asahina, S., Ishihara, T. & Yasumatsu, K. (Eds.), Iconographia Insectorum Japonicorum Colore naturali edita Volumen III. Hokuryu-kan Publishing, Tokyo, p. 289, 1 pl. [in Japanese]
Itô, Y., Miyagi, K. & Ota, H. (2000) Imminent extinction crisis among the endemic species of the forests of Yanbaru, Okinawa, Japan. Oryx, 34 (4), 305–316.
https://doi.org/10.1046/j.1365-3008.2000.00136.x
Kameyama, T., Nishi, M. & Nakamura, S. (2006) Insects from Ohta River, Hiroshima Prefecture, A result of survey in 2003. Miscellaneous Reports of the Hiwa Museum for Natural History, 47, 1–184. [in Japanese]
Kim, C.W. (1963) Hymenoptera of Korea. Humanities and Sciences (Natural Science), Korea University, 6, 243–374. [in Korean]
Kim, C.W. (1970) Illustrated Encyclopedia of Fauna and Flora of Korea, 11 (3). Samwha-Chulpansa, Seoul, 891 pp. [in Korean]
Kim, C.W., Lee, J.W., Park, J.S., Kim, B.J. & Park, J.C. (1994) Hymenoptera. In: Shin, Y.H. & Yoon, I.B. (Eds.), Check List of Insects from Korea. Kon-Kuk University Press, Seoul, pp. 216–269.
Kinota, K., Haneda, Y. & Tano, T. (2005) Aculeate wasps collected in Sapporo-shi, Otaru-shi and Ishikari-shi, Hokkaido. Tsunekibachi, 5, 25–31.
Kizaki, K. & Oshiro, I. (1977) Paleogeography of the Rytkyu [!] Islands. Marine Science Monthly, 9, 542–549. [in Japanese]
Kohl, F.F. (1880) Die Rauwespen Trirols nach ihrer horizontalen und verbreitung mit einen Anhange biologischen und kritischer Notizen. Z. Ferdinandeums Tirol Vorariberg, 3 (24), 97–242.
Kurokawa, H. (2005) Addition records of the Aculeata from Mt. Ashiwa-yama, Fukui. Tsunekibachi, 4, 76. [in Japanese]
Krombein, K.V. (1972) Monograph of the Madagascan Mutillidae (Hymenoptera). Part 1: Myrmillini. Mutillini and Smicromyrmini. Annales, Musée Royal de l’Afrique Centrale (Série in 8°), Sciences Zoologiques, 199, i–x + 1–61.
Krombein, K.V. (1978) Biosystematic studies of Ceylonese wasps III. Life history, nest and associates of Paraleptomenes mephitis (Cameron) (Hymenoptera: Eumenidae). Journal of the Kansas Entomological Society, 51 (4), 721–734.
Kubo, K. (2000) Order Hymenoptera. In: Iwase, K., Ohtsubo, H., Kubo, K., Shimura, T. & Watari, H. (Eds.), Insects of Mt. Enkai-zan and surrounding area. Kanagawa-Chûhô, 130, 287–343. [in Japanese]
Kuribayashi, S. (1973) Insects of Okinawa. Gakushu-Kenkyu-sha, Tokyo, 184 pp. [in Japanese]
Latreille, P.A. (1802) Histoire naturelle, générale et particulière des Crustacés et des Insectes. 3. F. Dufart, Paris, xii + 467 pp.
Lelej, A.S. (1976) Addition to the fauna of velvet ants (Hymenoptera, Mutillidae) of Mongolia. Insects of Mongolia, 4, 268–281. [in Russian]
Lelej, A.S. (1977a) Interesting data on Myrmosidae and Mutillidae (Hymenoptera) from the south of the Primorye territory. In: Lehr, P.A. (Ed.) Insects of the Russian Far East. Dalnauka, Vladivostok, pp. 88–95. [in Russian]
Lelej, A.S. (1977b) New data on the velvet-ant fauna of Mongolia (Hymenoptera, Mutillidae). Insects of Mongolia, 5, 526–536. [in Russian]
Lelej, A.S. (1978) Superfamily Mutilloidea. In: Tobias, V.I. (Ed.), Keys to the insects of the European part of the USSR. Vol. 3, pt 1. Nauka, Leningrad, pp. 71–84. [in Russian]
Lelej, A.S. (1981) Wasps of the families Myrmosidae and Mutillidae (Hymenoptera) from Eastern Siberia and the Far East. In: Lehr, P.A. (Ed.), Taxonomy of insects of the Far East. Dalnauka, Vladivostok, pp. 73–79. [in Russian].
Lelej, A.S. (1985) The velvet ants (Hymenoptera, Mutillidae) of the USSR and neighboring countries. Nauka, Leningrad, 268 pp. [in Russian]
Lelej, A.S. (1995a) A review of Palaearctic and Oriental species of genus Nemka Lelej with description of Oriental genus Mickelomyrme gen. n. (Hymenoptera, Mutillidae). Far Eastern Entomologist, 6, 1–20.
Lelej, A.S. (1995b) To the knowledge of East Asian species of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of four new genera and eight new species. Far Eastern Entomologist, 13, 1–28.
Lelej, A.S. (1996a) To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species. Far Eastern Entomologist, 30, 1–24.
Lelej, A.S. (1996b) Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics, 6 (1–2), 91–104.
https://doi.org/10.3759/tropics.6.91
Lelej, A.S. (1996c) A review of the East Asian species of Mickelomyrme Lelej, 1995 (Hymenoptera, Mutillidae). Entomofauna, 17 (15), 277–292.
Lelej, A.S. (2002) Catalogue of the Mutillidae (Hymenoptera) of the Palaearctic Region. Dalnauka, Vladivostok, 172 pp.
Lelej, A.S. (2005) Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok, 252 pp.
Lelej, A.S. (2007) Biogeography of mutillid wasps (Hymenoptera, Mutillidae). In: Rasnitsyn, A.P. & Gokhman, V.E. (Eds.), Studies on Hymenopterous Insects. Collection of Scientific Papers. KMK Scientific Press Ltd., Moscow, pp. 82–111. [in Russian]
Lelej, A.S. (2017) Family Mutillidae. In: Belokobylskij, S.A. & Lelej, A.S. (Eds.), Annotated Catalogue of the Hymenoptera of Russia. Vol. I. Symphyta and Apocrita: Aculeata. Proceedings of the Zoological Institute of the Russian Academy of Science, Supplement 6, pp. 152–158.
Lelej, A.S. & Brothers, D.J. (2008) The genus-group names of Mutillidae (Hymenoptera) and their type species, with a new genus, new name, new synonymies, new combinations and lectotypifications. Zootaxa, 1889, 1–79.
Lelej, A.S., Choi, J.Y. & Tripotin, P. (2001) Review of the mutillid wasps (Hymenoptera, Mutillidae) of Korea. Far Eastern Entomologist, 96, 1–15.
Lelej, A.S., Fateryga, A.V. & Ivanov, S.P. (2016) The velvet ants (Hymenoptera: Mutillidae) of the Crimean Peninsula. Far Eastern Entomologist, 314, 1–24.
Lelej, A.S., Gharali, B. & Lotfalizandeh, H. (2008) New records of velvet ants (Hymenoptera: Mutillidae) from the west of Iran. Far Eastern Entomologist, 191, 1–7.
Lelej, A.S. & van Harten, A. (2006) A review of the Mutillidae (Hymenoptera) of Yemen. Zootaxa, 1226 (1), 1–50.
https://doi.org/10.11646/zootaxa.1226.1.1
Lelej, A.S. & van Harten, A. (2014) Order Hymenoptera, family Mutillidae. Genus Physetopoda Schuster, 1949. In: van Harten, A. (Ed.), Arthropod fauna of the UAE. Vol. 5. Dar AI Ummah, Abu Dhabi, pp. 505–520.
Lelej, A.S. & Kabakov, O.N. (1980) On Bradynobaenidae and Mutillidae (Hymenoptera) of Afghanistan. Entomologicheskoe Obozrenie, 59(1), 181–196. [in Russian, English translation (1981) in Entomological Review, 59 (1), 141–156]
Lelej, A.S. & Nemkov, P.G. (1997) Phylogeny, evolution and classification of Mutillidae (Hymenoptera). Far Eastern Entomologist, 46, 1–24.
Lelej, A.S., Proshchalykin, M.Yu. & Loktionov, V.M. (2014) To the knowledge of velvet ants (Hymenoptera, Mutillidae) of East Siberia. Euroasian Entomological Journal, 13 (4), 388–394. [in Russian]
Lelej, A.S. & Shlyakhtenok, S. (2015) The velvet ants (Hymenoptera: Mutillidae) of Belarus, with analysis of the fauna of eastern European countries. Far Eastern Entomologist, 297, 1–15.
Lelej, A.S., Ullah, M. & Mahmood, K. (2007) Additions to the knowledge of the Mutillidae (Hymenoptera) of Pakistan. Zootaxa, 1444 (1), 53–60.
https://doi.org/10.11646/zootaxa.1444.1.4
Lelej, A.S. & Yamane, Sk. (1992) Interesting velvet ants (Hymenoptera, Mutillidae) from Japan and Korea. Japanese Journal of Entomology, 60 (3), 625–632.
Lepeletier de Saint-Fargeau, A. (1845) Histoire Naturelle des Insectes. Hyménoptères. Tome Troisième. Roret, Paris, [4] + 646 pp.
Lo Cascio, P. (2015) Worldwide checklist of the island mutillid wasps (Hymenoptera Mutillidae). Biodiversity Journal, 6 (2), 529–592.
Lo Cascio, P. & Grita, F. (2014) A new species of Smicromyrme from Israel (Hymenoptera: Mutillidae). Fragmenta entomologica, 46 (1–2), 125–128.
https://doi.org/10.4081/fe.2014.76
Magretti, P. (1892) Viaggio di Leonardo Fea in Birmania e regioni vicine XLIII. Imenotteri. Parte prima. Mutillidei, Scoliidei, Tifiidei, Tinnidei colla descrizione di parecchie nuove specie. Annali del Museo Civico di Storia Naturale di Genova, 2 (12), 197–266, 1 pl.
Marikovskaya, T.P. (1968) New data on biology of some colonial bees (Hymenoptera, Megachilidae) in south-east Kazakhstan. Entomologicheskoye obozreniye, 47 (4), 796–805. [in Russian]
Maruyama, S., Isozaki, Y., Kimura, G. & Terabayashi, M. (1997) Paleogeographic maps of the Japanese Islands: Plate tectonic synthesis from 750 Ma to the present. Island Arc, 6, 121–142.
https://doi.org/10.1111/j.1440-1738.1997.tb00043.x
Matsuda, S. & Nakamura, S. (1999) The Insects of the basin of the Gouno River, Hiroshima and Shimane Prefecture. Miscellaneous Reports of the Hiwa Museum for Natural History, 37, 39–176. [in Japanese]
Matsui, M., Ito, H., Shimada, T., Ota, H., Saidapur, S.K., Khonsue, W., Tanaka-Ueno, T. & Wu, G.-F. (2005a) Taxonomic relationships within the Pan-Oriental narrow-mouth toad Microhyla ornata as revealed by mtDNA analysis (Amphibia, Anura, Microhylidae). Zoological Science, 22, 489–495.
https://doi.org/10.2108/zsj.22.489
Matsui, M., Shimada, T., Ota, H. & Tanaka-Ueno, T. (2005b) Multiple invasions of the Ryukyu Archipelago by Oriental frogs of the subgenus Odorrana with phylogenetic reassessment of the related subgenera of the genus Rana. Molecular Phylogenetics and Evolution, 37, 733–742.
https://doi.org/10.1016/j.ympev.2005.04.030
Matsumura, S. & Uchida, T. (1926) Die Hymenopteren-Fauna von den Riukiu Inseln. Insecta Matsumurana, 1, 32–52, 63–77 + 3 pls.
Mickel, C.E. (1933) The Mutillidae of Formosa. Annals of the Entomological Society of America, 26 (2), 381–423.
https://doi.org/10.1093/aesa/26.2.381
Mickel, C.E. (1934) Mutillidae of the Philippine Islands. Philippine Journal of Science, 54 (1), 91–219, 1 pl.
Mickel, C.E. (1935) The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London, 83 (2), 177–312.
https://doi.org/10.1111/j.1365-2311.1935.tb01207.x
Mickel, C.E. (1936) A new species of Smicromyrme from Japan (Mutillidae, Hymenoptera). Mushi, 9 (1), 52–54.
Mickel, C.E. (1937) New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands. The Annals and Magazine of Natural History, Series 10, 19 (112), 441–456.
https://doi.org/10.1080/00222933708655284
Ministry of Environment (2015) Red Data Book 2014. Threatened Wildlife of Japan. Gyosei, Tokyo, xxxix + 509 pp., 8 pls.
Motokawa, M. (2017) “Land Emergence” and “Elevation Shift” affect diversification: A new perspective toward understanding the high species diversity of terrestrial animals in Japan. In: Motokawa, M. & Kajihara, H. (Eds.), Species Diversity of Animals in Japan. Springer Japan, Tokyo, pp. 3–23.
https://doi.org/10.1007/978-4-431-56432-4_1
Munakata, M. (1987) Records of bees and wasps from Okushiri-tô Island. Hymenopterists’ Communication, 27, 35–39. [in Japanese]
Murota, T. & Nozaka, C. (1987) Records of wasps from Yaeyama Islands. Hymenopterists’ Communication, 27, 106–116. [in Japanese]
Muskovits, J. & Lelej, A.S. (2010) Smicromyrme pilisensis sp. n. from Hungary. Folia Entomologica Hungaria, 71, 57–63.
Nagase, H. (2004) Hymenoptera (excl. Formicidae). In: Kanagawa Konchû Danwakai (Ed.), Insect fauna of Kanagawa. III. Kanagawa Konchû Danwakai, Odawara, pp. 1241–1326. [in Japanese]
Nagase, H., Harada, K., Ito, H. & Yago, M. (2014), Catalogue of the Masayo Kato insect collection, the University Museum, the University of Tokyo Part I. Hymenoptera. The University Museum, The University of Tokyo Material Reports, 104, 5–189.
Nagy, C.G. (1966) Ein neues Mutilliden-Subgenus aus Rumänien (Hymenoptera). Reichenbachia, 6 (13–34), 113–117.
Nakamura, S. & Haneda, Y. (1997) Distributional notes on Wasps (Apocrita: Hymenoptera) from Hiroshima Prefecture. Miscellaneous Reports of the Hiwa Museum for Natural History, 35, 31–48. [in Japanese]
Nakamura, S. & Haneda, Y. (2000) Distributional notes on Wasps and Bees (Apocrita: Hymenoptera) from Hiroshima Prefecture (4). Miscellaneous Reports of the Hiwa Museum for Natural History, 39, 117–136. [in Japanese]
Nakamura, S., Kameyama, T. & Katayama, S. (2001) The Insects in the Riverside of Gono-kawa River. Bulletin of the Hoshizaki Green Foundation, 5, 17–114. [in Japanese]
Nakamura, S., Takayama, T. & Hamaguchi, K. (2000) Insects from Ashida River, Hiroshima Prefecture, A result of survey in 1997. Journal of the Hiba Society of Natural History, 196, 1–144. [in Japanese]
Negoro, H., Haneda, Y. & Tano, T. (2005) The Aculeata of Toyama Prefecture. Tsunekibachi, 4, 31–52. [in Japanese]
Nurse, C.G. (1903) New species of Indian Aculeate Hymenoptera. The Annals and Magazine of Natural History, Series 7, 11 (64), 393–403 + 511–526 + 528–549.
https://doi.org/10.1080/00222930308678789
Ogawa, H., Nosaka, M., Hashii, N., Yokoyama, M. & Tsurusaki, N. (2012) Records of faunal survey of insects of Tottori Sand Dunes in 2011 with comments to the fauna of “Sakyu Oasis”. Natural History Research of San’in, 7, 31–40. [in Japanese]
Ohkusa, S. (2006) Hymenoptera collected from Gifu Prefecture. Tsunekibachi, 9, 16–28. [in Japanese]
Ohkusa, S. (2007) Hymenoptera collected from North-West at Aichi Prefecture. Tsunekibachi, 12, 26–38. [in Japanese]
Ohkusa, S. & Tajima, T. (2008) Hymenoptera collected from North-West at Aichi Prefecture. Tsunekibachi, 13, 63–81. [in Japanese]
Ohkusa, S. & Tajima, T. (2010) Hymenoptera of eastern part of Shiga Prefecture. Tsunekibachi, 17, 1–20. [in Japanese]
Okayasu, J. (2017) Two new species of the genus Sinotilla Lelej, 1995 (Hymenoptera: Mutillidae), with notes on taxonomic characters. Zootaxa, 4294 (2), 151–169.
https://doi.org/10.11646/zootaxa.4294.2.1
Okayasu, J. (2018) Taxonomic review of Chin-wen Chen’s species described in the genus Smicromyrme (Hymenoptera: Mutillidae: Smicromyrmini). Acta Entomologica Musei Nationalis Pragae, 58 (2), 479–494.
https://doi.org/10.2478/aemnp-2018-0036
Okayasu, J., Williams, K.A. & Lelej, A.S. (2018) A remarkable new species of the genus Sinotilla Lelej (Hymenoptera: Mutillidae: Smicromyrmini) from Taiwan and an overview of color diversity in East Asian mutillid females. Zootaxa, 4446 (3), 301–324.
https://doi.org/10.11646/zootaxa.4446.3.1
Olivier, A.G. (1812) Oxybèle. Encyclopédie Methodique. Histoire Naturelle. Insectes, 8, 592–597.
Ono, H. & Kawashima, I. (2013) Host records on two species of mutillid wasps (Hymenoptera, Mutillidae) in Japan. Tsunekibachi, 23, 41–42. [in Japanese]
Ota, H. (1998) Geographic patterns of endemism and speciation in amphibians and reptiles of the Ryukyu Archipelago, Japan, with special reference to their Paleogeographical implications. Researches on Population Ecology, 40 (2), 189–204.
https://doi.org/10.1007/BF02763404
Ovchinnikov, S.V. (2002) Materials to the fauna of velvet ants (Hymenoptera, Mutillidae) of Kyrghizia. Entomological Investigations in Kyrgystan, 22, 79–104. [in Russian]
Pagden, H.T. (1934) New species of Mutillidae (Hymenoptera, Vespoidea) from the Malay Peninsula. Journal of the Federated Malay States Museums, 17, 419–457.
Pagden, H.T. (1949) Descriptions and records of Austro-Malaysian Methocidae and Mutillidae (Hymenoptera). Transactions of the Royal Entomological Society of London, 100 (8), 192–231.
https://doi.org/10.1111/j.1365-2311.1949.tb01431.x
Pagliano, G., Palmerini, M.M. & Strumia, F. (2018) Una nuova specie di Astomyrme Schwartz, 1984 del Sud Italia (Insecta: Hymenoptera: Mutillidae). Quaderno di Studi e Notizie di Storia Naturale della Romagna, 48, 139–146.
Pagliano, G. & Strumia, F. (2007) Contributo alla conoscenza dei Mutillidae italiani (Hymenoptera, Scolioidea). Bollettino del Museo Regionale di Scienze Naturali, Torino, 24 (1), 25–110.
Pagliano, G. & Strumia, F. (2013) Smicromyrme corriasi n. sp. from Sardinia and new data on some Italian Smicromyrme and Stenomutilla (Hymenoptera, Mutillidae). Naturalista Valtellinese, 24, 55–66.
Pallas, P.S. (1773) Reisen durch verschiedene Provinzen des Russischen Reichs. Zweiter Theil. Zweites Buch vom Jahr 1771. Kayserlichen Academie der Wissenschaften, St. Petersburg, [12] + 744 pp.
Pérez, J. (1905) Hyménoptères recueillis dans le Japon central, par M. Harmand, ministre plénipotentiaire de France à Tokio. Troisième partie. Pompilides, Pemphrédonides, Sphégides et Crabronides. Bulletin de Muséum d’Histoire naturelle, 3, 148–158.
Pitts, J.P. & Manley, D.G. (2004) A revision of Lomachaeta Mickel, with a new species of Smicromutilla Mickel (Hymenoptera: Mutillidae). Zootaxa, 474 (1), 1–27.
https://doi.org/10.11646/zootaxa.474.1.1
Radoszkowski, O. (1865) Les mutilles russes. Bulletin de la Société impériale des naturalistes de Moscou, 38, 422–467, tabs. 7–9.
Sakagami, S.F., Miyanaga, R. & Maeta, T. (1994) Discovery of a eusocial halictine bee, Lasioglossum (Evylaeus) subtropicum sp. nov. from Iriomote Is., southernmost Japan, with a morphometric comparison of castes in some social halictines (Hymenoptera, Halictidae). Bulletin of the Faculty of Agriculture, Shimane University, 28, 5–21.
Sato, T. & Tsurusaki, N. (2010) Preliminary report of insect fauna occurring in the Tottori Sand Dunes. Bulletin of the Tottori Prefectural Museum, 47, 45–81. [in Japanese]
Schulz, W.A. (1906) Spolia Hymenopterologica. Albert Pape, Paderborn, 356 pp., 1 pl.
Schuster, R.M. (1947) A revision of the Sphaerophthalmine Mutillidae of America north of Mexico. Annales of the Entomological Society of America, 39 (4), 692–703.
Schuster, R.M. (1949) Contribution towards a monograph of the Mutillidae of the Neotropical region. III. A key to the subfamilies represented and descriptions of several new genera (Hymenoptera). Entomologica Americana, New Series, 29 (3–4), 59–140.
Schwartz, F. (1984) Description d’un genre nouveau, disjoint de Smicromyrme, et d’une espèce nouvelle du sud de la France (Hymenoptera, Mutillidae). L’Entomologiste, 40, 21–25.
Serkova, L.G. (1962) To the natural history of desert pasture pest leaf beetle (Theone silphoides). Proceedings of the Institute of Plant Protection of Kazakh Soviet Socialist Republic, 7, 83–108. [in Russian]
Shintani, T. (2003) Distributional information on insects from Sera-plateau, Hiroshima Prefecture (1). Journal of the Hiba Society of Natural History, 208, 13–19, 1 pl. [in Japanese]
Shorthouse, D.P. (2010) SimpleMappr, an online tool to produce publication-quality point maps. Available from: http://www.simplemappr.net (accessed 25 April 2019)
Sichel, J. & Radoszkowski, O. (1869–1870) Essai d’une monographie des Mutilles de l’Ancion Continent. Horae Societatis Entomologicae Rossicae, 1869 & 1870, 6 (3) & 6 (4), 139–172 & 173–309, tabs. 6–11.
Skorikov, A.S. (1935) Zur Mutilliden-Fauna Zentralasiens. Trudy Tadzhikskoi Basy Akademii Nauk SSSR, 5, 257–349 + 7 pls. [in Russian]
Smith, F. (1855) Catalogue of Hymenopterous Insects in the Collection of the British Museum. Part III. Mutillidae and Pompilidae. Taylor and Francis, London, 206 pp., 5 pls.
Smith, F. (1857–1858) Catalogue of the hymenopterous insects collected at Sarawak, Borneo; Mount Ophir, Malacca; and at Singapore, by A.R. Wallace. Journal of the Proceedings of the Linnean Society, Zoology, 2, 1857 & 1858, 42–88, pls. 1–2 & 89–130.
https://doi.org/10.1111/j.1096-3642.1857.tb01759.x
Smith, F. (1860 [1861]) Catalogue of hymenopterous insects collected by Mr. A.R. Wallace in the islands of Bachian, Kaisaa, Amboyna, Gilolo, and at Dory in New Guinea. Journal of the Proceedings of the Linnean Society, Zoology, 5, 93–143.
https://doi.org/10.1111/j.1096-3642.1860.tb01022.x
Smith, F. (1873) Description of Aculeate Hymenoptera of Japan, collected by Mr. George Lewis at Nagasaki and Hiogo. The Transactions of the Entomological Society of London, 1873, 181–206.
https://doi.org/10.1111/j.1365-2311.1873.tb00641.x
Smith, F. (1879) Descriptions of new species of Hymenoptera in the collection of the British Museum. Taylor and Francis, London, XXI + 240 pp.
Snodgrass, R.E. (1941) The male genitalia of Hymenoptera. Smithsonian Miscellaneous Collections, 99 (14), 1–86, 33 pls.
Sonan, J. (1931) Some wasps and bees of Hokoto (Pescadores). Transactions of the Natural History Society of Formosa, 21 (112), 6–8. [in Japanese]
Suárez, F.J. (1959) Especies nuevas o poco conocidas de la fauna Mediterránea II nota. Eos, Madrid, 35 (3), 283–300, 5 pls.
Takahashi, H. (1996) Records of Mutillidae. Gekkan-Mushi, 310, 28–29. [in Japanese]
Takahashi, H. & Matsumoto, Y. (1999) Some records of Hymenoptera from Izu Islands, Tokyo, Japan. Gensei, 74, 23–26. [in Japanese]
Tano, T., Kurokawa, H., Murota, T. & Nozaka, C. (2003) The Hymenoptera collected at Tsushima, Nagasaki Prefecture in July of 2003. Tsunekibachi, 1, 23–40. [in Japanese]
Templado, J. (1958) Smicromyrme viduata Pall. parasito de Bembix mediterranea Handl. Graellsia, 16 (1–3), 45–47.
Terayama, M. (2005) Taxonomic guide to the Japanese Aculeate wasps, 2. Family Mutillidae. Tsunekibachi, 4, 1–13. [in Japanese]
Terayama, M. (2019) Record of a female velvet ant of the genus Ephucilla collected in Okinawa Island. Gekkan-Mushi, 580, 48. [in Japanese]
Terayama, M., Suda, H., Murota, T., Haneda, Y. & Tano, T. (2011) The velvet ants (Mutillidae) of Japan. Gekkan-Mushi, 481, 12–25. [in Japanese]
Terayama, M., Suda, H., Murota, T. & Tano, T. (2016) Family Mutillidae. In: Terayama, M. & Suda, H. (Eds.), A guide to the Aculeate wasps of Japan. Tokai University Press, Kanagawa, pp. 161–174. [in Japanese]
Thomson, C.G. (1870) Öfversigt af Sveriges rofsteklar. Opuscula Entomologica, Lund, 1 (2), 202–251.
Toda, M., Nishida, M., Tu, M.-C., Hikida, T. & Ota, H. (1999) Genetic variation, phylogeny and biogeography of the pit vipers of the genus Trimeresurus sensu lato (Reptilia: Viperidae) in the subtropical East Asian island. In: Ota, H. (Ed.), Tropical island herpetofauna: origin, current diversity, and conservation. Elsevier, Amsterdam, pp. 249–270.
Tormos, J., Asis, J.D., Beneitez, A. & Gayubo, S.F. (2009) Description of the mature larva of the sand wasp Bembix bidentata and its parasitoids (Hymenoptera: Crabronidae, Chrysididae, Mutillidae). Florida Entomologist, 92 (1), 43–53.
http://doi.org/10.1653/024.092.0108
Tormos, J., Asis, J.D. & Gayubo, S.F. (2003) Description of the mature larva of Nemka viduata (Pallas) (Hymenoptera: Multillidae: Mutillinae), a parasitoid of Stizus continuus (Klug) (Hymenoptera: Crabronidae: Bemibicinae). Journal of Entomological Science, 38 (4), 502–510.
https://doi.org/10.18474/0749-8004-38.4.502
Tonnoir, A.L. (1930) Notes on Indian Blepharocerid larvae and pupae with remarks on the morphology of Blepharocerid larvae and pupae in general. Records of the Indian Museum, 32, 161–214.
Tsuneki, K. (1965). The biology of east-Asiatic Cerceris (Hym., Sphecidae) with special reference to the peculiar social relationships and return to the nest in Cerceris hortivaga Kohl. Etizenia, 9, 1–46.
Tsuneki, K. (1972a) Studies on the Mutillidae of Japan (Hymenoptera). Etizenia, 61, 1–26.
Tsuneki, K. (1972b) Two new species of Mutillidae from Borneo (Hym.). The Life Study, 16 (1–2), 17–21.
Tsuneki, K. (1972c) Mutillidae collected in Formosa in 1966 and 68 (Hymenoptera). Etizenia, 64, 1–25.
Tsuneki, K. (1973) New and the first recorded species and subspecies of Sphecidae and Mutillidae from Japan, with taxonomic notes on some species (Hymenoptera). Etizenia, 65, 1–28.
Tsuneki, K. (1993a) On some Taiwanese Mutillidae, collected in 1976 by Mr. T. Murota, with description of new taxa (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 41, 39–50.
Tsuneki, K. (1993b) A new and an unrecorded species of Mutillidae from Is. Okinawa, the Ryukyus (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 41, 51–55.
Tsuneki, K., Nozaka, C., Tano, T., Kurokawa, H. & Murota, T. (1993) Mutillidae recently collected in the Philippines (Hymenoptera). Special Publications of the Japan Hymenopterists Association, 41, 1–38.
Tsurusaki, N., Hayashi, M., Miyanaga, R., Ichisawa, K. & Kawakami, Y. (2012) Catalogue of the insects of Tottori Sand Dunes, Honshu, Japan. Natural History Research of San’in, 7, 47–82. [in Japanese]
Turner, R.E. (1916) On Mutillidae parasitic on Glossina morsitans. Bulletin of Entomological Research, London, 7, 93–95.
https://doi.org/10.1017/s0007485300017119
Turrisi, G.F. & Foucart, A. (2008) Description of a gynandromorphic Myrmilla calva (Villiers 1789), with a summary on gynandromorphism phenomenon within Mutillidae (Hymenoptera: Scolioidea). Linzer Biologische Beiträge, 40 (2), 1873–1880.
Williams, K.A., Brothers, D.J. & Pitts, J.P. (2011) New species of Tobantilla Casal, 1965 and a new genus and species, Gogoltilla chichikovi gen. et sp. nov., from Argentina (Hymenoptera: Mutillidae). Zootaxa, 3064 (1), 41–68.
https://doi.org/10.11646/zootaxa.3064.1.2
Williams, K.A., Cambra, R.A., Waldren, G.C., Quintero A., D. & Pitts, J.P. (2019a) Silvorientilla, gen. nov. a new Neotropical velvet ant genus (Hymenoptera: Mutillidae). Folia Entomológica Mexicana, Nueva Serie, 4 (3), 91−109.
Williams, K.A., Lelej, A.S., Okayasu, J., Borkent, C.J., Malee, R., Thoawan, K. & Thaochan, N. (2019b) The female velvet ants (aka modkhong) of southern Thailand (Hymenoptera: Mutillidae), with a key to the genera of southeast Asia. Zootaxa, 4602 (1), 1–69.
https://doi.org/10.11646/zootaxa.4602.1.1
Yamada, H. (1990) Mutillidae, Chrysididae, Sphecidae and Megachilidae of Aichi Prefecture. In: Natural Environment Division of Aichi Prefecture (Ed.), The Insects of Aichi (the first volume). Aichi Prefecture Government, Aichi, pp. 480–487. [in Japanese]
Yamada, M. (2001) Aculeate Hymenoptera fauna of Aomori Prefecture, the northernmost Honshu, Japan. 10. Distributional records of mutillid, tephiid (!) and scoliid wasps. Journal of the natural history of Aomori, 6, 57–60. [in Japanese]
Yamamoto, E. & Haneda, Y. (2007) Aculeata collected in Odamiyama and the surrounding area (supplement 3). Shikokukogera, 4, 15–27. [in Japanese]
Yamane, Sk. (1999) Family Mutillidae. In: Yamane, Sk., Ikudome, S. & Terayama, M. (Eds.), Identification guide to the Aculeata of the Nansei Islands, Japan. Hokkaido University Press, Sapporo, pp. 369–381.
Yano, M. (1932) Mutillidae. In: Uchida, S. (Ed.), Iconographia Insectorum Japonicorum Editio Prima. Hokuryukan, Tokyo, pp. 325–327, 1 pl.
Yasumatsu, K. (1934) Les Hyménoptères de l’ile Yakushima. Mushi, 7 (2), 61–67.
Yasumatsu, K. (1937) Hymenoptera of Tsushima (first report). Journal of Natural History Society in Fukuoka, 2 (2), 59–74. [in Japanese]
Yasumatsu, K. (1938a) Les Mutillides de Coree et de la Mandschourie du Sud. Transactions of the Natural History Society of Formosa, 28 (175), 83–87.
Yasumatsu, K. (1938b) Beitrage zur Synonymie einiger Hymenopterenarten von den Ryukyu-Inseln. Transactions of the Natural History Society of Formosa, 28 (183), 446–447.
Yasumatsu, K. (1938c) Family Mutillidae. In: Esaki, T., Hori, H. & Yasumatsu, K. (Eds.), Insectorum Japonicorum Illastratio Iconographica Coloribus ad Naturam Depicta. Sanseidô, Tokyo, p. 351. [in Japanese]
Yasumatsu, K. (1951) Mutillidae of Shansi, North China (Hymenoptera). Mushi, 22 (12), 69–72.
Yoshida, M. (1989) Family Mutillidae. In: Entomological Laboratory, Faculty of Agriculture, Kyushu University & Japan Wildlife Research Center (Eds.), A Check List of Japanese Insects. Entomological Laboratory, Faculty of Agriculture, Kyushu University, Fukuoka, pp. 650–651. [in Japanese]
Zavattari, E. (1913) H. Sauter’s Formosa-Ausbeute. Mutillidae (Hym.). Archiv für Naturgeschichte, 79A (3), 19–42.
https://doi.org/10.5962/bhl.part.7709
Zavattari, E. (1914) Mutille Austro-Malesi. Bolletino della Societa Entomologica Italiana, 45, 61–114.
Zimmermann, K. (1931) Studien über individuelle und geographische variabilität paläarktischer Polistes und verwandter Vespiden. Zeitschrift für Morphogie und Ökologie der Tiere, 22, 173–230.
https://doi.org/10.1007/BF00417968
Zhou, H.-T., Lelej, A.S. & Liu, J.-X. (2018) New records of velvet ants (Hymenoptera: Mutillidae) from China with taxonomic notes. Far Eastern Entomologist, 354, 23–28.
https://doi.org/10.25221/fee.354.4