Abstract
The Barsine hypoprepioides (Walker, 1862) species-group is reviewed. Forty seven species and one subspecies belong to the species-group. Among them, fifteen species and one subspecies are new for science and described in the present paper: B. amoenissima sp. n. (Myanmar), B. selene sp. n. (Thailand and Laos), B. cao sp. n. (Thailand, Laos and Vietnam), B. speideli sp. n. (SE China, Thailand, Laos, Cambodia and Vietnam), B. mesomene sp. n. (NE India, Myanmar, China, Thailand, Laos and Vietnam), B. pandeia sp. n. (Vietnam), B. karenkensis wushipheri ssp. n. (S Taiwan), B. euryphaessa sp. n. (Vietnam), B. ivanovamariae sp. n. (Vietnam and SE China), B. eos sp. n. (Thailand), B. mene sp. n. (Thailand), B. syntypicoida sp. n. (Malaysia and Indonesia), B. dubatolovi sp. n. (The Philippines), B. visaya sp. n. (The Philippines), B. nemea sp. n. (The Philippines) and B. kishidai sp. n. (The Philippines). Eight new combinations are established: B. parameia (Rothschild, 1913), comb. n. B. takamukui (Matsumura, 1927), comb. n., B. karenkensis (Matsumura, 1930), comb. n., B. rhipiptera (Wileman & West, 1928), comb. n., B. chi (Roepke, 1946), comb. nov., B. salakia (Schaus, 1922), comb. n., B. marginis (Fang, 1991), comb. nov. and B. dentata (Wileman, 1910), comb. nov. The lectotype is designated for Lithosia hypoprepioides Walker, 1862. Adults, male and female genitalia of all species are illustrated.
References
Bae, Y.S., Bayarsaikhan, U. & Kimsun, C. (2016) Biodiversity of Cambodia. Tiger Moths (Insecta, Lepidoptera). National Institute of Biological Resources, Incheon, 303 pp.
Bayarsaikhan, U., Lee, D.J. & Bae, Y.S. (2018) A review of Barsine Walker, 1854 (Lepidoptera: Erebidae, Arctiinae) in Cambodia, with a new record. Journal of Asia-Pacific Biodiversity, 11 (3), 377–385.
https://doi.org/10.1016/j.japb.2018.05.003
Bucsek, K. (2012) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malaya Peninsula-Malaysia. Institute of Zoology SAS, Bratislava, 170 pp.
Bucsek, K. (2014) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malaya Peninsula-Malaysia (Supplementum). Institute of Zoology SAS, Bratislava, 45 pp.
Butler, A.G. (1877) Descriptions of new Species of Heterocera from Japan.—Part I. Sphinges and Bombyces. The Annals and magazine of natural history, 320 (119), 93–404.
https://doi.org/10.1080/00222937708682255
Černý, K. (2016) A contribution to the knowledge of the Miltochrista-Lyclene genus group in South East Asia (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Nachrichten des entomologischen Vereins Apollo, 37 (2–3), 93–107.
Černý, K. & Pinratana, A. (2009) Moths of Thailand. Vol. 6. Arctiidae. Brothers of Saint Gabriel in Thailand, Bangkok, 283 pp.
Daniel, F. (1951) Beiträge zur Kenntniss der Arctiidae Ostasiens unter besonderer Berücksichtigung der Ausbeuten von Dr. h. c. H. Höne aus diesem Gebiet (Lep.-Het.). III. Teil: Lithosiinae. Bonner zoologische Beiträge, 2 (3–4), 291–327. [in German]
Daniel, F. (1952) Beiträge zur Kenntniss der Arctiidae Ostasiens unter besonderer Berücksichtigung der Ausbeuten von Dr. h. c. H. Höne aus diesem Gebiet (Lep.-Het.). III. Teil: Lithosiinae. Bonner zoologische Beiträge, 3 (1–2), 75–90. [in German]
Daniel, F. (1955) Beiträge zur Kenntniss der Arctiidae Ostasiens unter besonderer Berücksichtigung der Ausbeuten von Dr. h. c. H. Höne aus diesem Gebiet (Lep.-Het.). IV. Teil: Nachträge. Bonner zoologische Beiträge, 6 (1–4), 132–145. [in German]
Dubatolov, V.V. & Bucsek, K. (2013) New species of lichen-moths from South-East Asia (Lepidoptera, Noctuoidea, Lithosiini). Tinea, 22 (4), 279–291.
Dubatolov, V.V., Kishida, Y. & Wang, M. (2012) New records of lichen-moth from the Nanling Mts., Guangdong, South China, with descriptions of new genera and species (Lepidoptera, Arctiidae: Lithosiinae). Tinea, 22 (1), 25–52.
Fang, C. (1991) Studies of the genus Miltochrista of China (Lepidoptera: Arctiidae: Lithosiinae). Sinozoologia, 8, 383–397. [in Chinese]
Fang, C. (2000) Lepidoptera. Arctiidae. Fauna Sinica (Insecta). 19. Science Press, Beijing, 590 pp. [in Chinese]
Felder, C. (1861) Lepidopterorum Amboinensium. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe, 1861 (1–5), 26–44. [in Latin]
Fu, C.M. & Tzuoo, H.R. (2004) Moths of Anmashan. Part 2. Taichung Nature Research Society, Taichung, 215 pp., pls 37–60.
Hampson, G.F. (1894) The Fauna of British India, including Ceylon and Burma.Moths. Vol. 2. Taylor and Francis, London, 609 pp.
Hampson, G.F. (1898) The moths of India. Supplementary paper to the volumes in “The fauna of British India”. Part I-II. Journal of the Bombay Natural History Society, 11 (3), 438–662.
Hampson, G.F. (1900) Catalogue of the Arctiadae (Nolinae, Lithosianae) in the British Museum. Catalogue of the Lepidoptera Phalaenae in the British Museum, 2, 1–590.
Holloway, J.D. (1982) Taxonomic appendix. In: Barlow, H.S. (Ed.), An introduction to the moths of South East Asia. The author, Kuala Lumpur, pp. 176–271.
Holloway, J.D. (2001) The moths of Borneo: family Arctiidae, subfamily Lithosiinae. Malayan Nature Journal, 7, 279–486.
Horsfield, T. & Moore, F. (1858 [1860]) A catalogue of the lepidopterous insects in the Museum of Natural History. East-India House, 2, 1–440.
Huang, S.Y., Ji, S.Q., Wang, M. & Fan, X.L. (2018) Barsine paraprominens sp. nov. (Lepidoptera, Erebidae, Arctiinae, Lithosiini) from China, with notes on four new records. Zootaxa, 4457 (1), 189–196.
https://doi.org/10.11646/zootaxa.4457.1.11
Inoue, H. (1980) Synonymic Notes on Some Lithosiinae, Arctiidae, from Japan and Taiwan, with Descriptions of Three New Species: Lepidoptera. Tyô to Ga, 30 (3, 4), 159–171.
Joshi, R., Singh, N. & Volynkin, A.V. (2018) New data on the genus Barsine Walker, 1854 from India, with description of a new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini). Zootaxa, 4425 (3), 456–470.
https://doi.org/10.11646/zootaxa.4425.3.2
Kaleka, A.P.S. (2003) Revival of genus Barsine Walker (Lithosiinae: Arctiidae: Lepidoptera) along with description of two new species from India. Journal of Entomological Research, 27 (2), 93–103
Kirti, J.S. & Singh N. (2016) Arctiid Moths of India. Vol. 2. Nature Books India, New Delhi, 214 pp.
Kishida, Y. (1978) The illustrations of moths from Taiwan, (12–16). Gekkan-Mushi, 90, 27–28. [in Japanese]
Kishida, Y. (2011) Lithosiinae. In: Wang, M. & Kishida, Y. (Eds.), Moths of Guangdong Nanling National Nature Reserve. Goecke & Evers, Keltern, pp. 216–223.
Leech, J.H. (1899) Lepidoptera Heterocera from Northern China, Japan and Corea. Part II. Transactions of the Entomological Society of London, 1899 (1), 99–219.
https://doi.org/10.1111/j.1365-2311.1899.tb03304.x
Matsumura, S. (1927) New species and subspecies of moths from the Japanese Empire. Journal of the College of Agriculture, Hokkaido Imperial University, 19 (1), 1–91.
Matsumura, S. (1930) New species and forms of Arctiidae from Japan. Insecta Matsumurana, 5, 31–40.
Moore, F. (1878) A revision of certain genera of European and Asiatic Lithosiinae, with characters of new genera and species. Proceedings of the Scientific Meetings of the Zoological Society of London, 1878, 3–37, pls. 1–3.
https://doi.org/10.1111/j.1469-7998.1878.tb07927.x
Roepke, W. (1946) The Lithosiids, collected by Dr. L.J. Toxopeus in Central Celebes, with remarsk on some allied species. Tijdschrift voor entomologie, 87, 77–91.
Rothschild, W. (1913) New Lithosianae. Novitates Zoologicae, 20 (1), 192–226.
Schaus, W. (1922) New species of Lithosiidae from the Oriental Region (Lepidoptera). Insecutor inscitiae menstruus, 10 (1–3), 23–37.
Swinhoe, Ch. (1906) Eastern and African Heterocera. The annals and magazine of natural history: zoology, botany, and geology, 17 (7), 540–556.
https://doi.org/10.1080/00222930608678909
Volynkin, A.V. (2018) Four new species of the genus Barsine Walker, 1854 (Lepidoptera: Erebidae, Arctiinae) from Oriental Region. Far Eastern Entomologist, 358, 1–18.
https://doi.org/10.25221/fee.358.1
Volynkin, A.V. & Černý, K. (2016a) Barsine amaculata, a new species from Vietnam (Lepidoptera, Erebidae, Arctiinae). Zootaxa, 4200 (2), 345–350.
https://doi.org/10.11646/zootaxa.4200.2.11
Volynkin, A.V. & Černý, K. (2016b) A review of the Barsine flammealis species˗complex with description of a new species from Himalaya (Lepidoptera, Erebidae, Arctiinae). Biological Bulletin of Bogdan Chmelnitskiu Melitopol State Pedagogical University, 6 (3), 303–310.
https:// doi.org/10.15421/201699
Volynkin, A.V. & Černý, K. (2016c) Barsine deliciosa, a new species from China (Lepidoptera, Erebidae, Arctiinae). Zootaxa, 4200 (1), 181–191.
https://doi.org/10.11646/zootaxa.4200.1.9
Volynkin, A.V. & Černý, K. (2017a) A review of the Barsine obsoleta species˗group with description of a new species from Vietnam. Zootaxa, 4254 (2), 188–200.
https://doi.org/10.11646/zootaxa.4254.2.2
Volynkin, A.V. & Černý, K. (2017b) On the taxonomy of the Barsine delineata species˗complex (Lepidoptera, Erebidae, Arctiinae). Zootaxa, 4258 (2), 138–144.
https://doi.org/10.11646/zootaxa.4258.2.3
Volynkin, A.V. & Černý, K. (2017c) Revision of the Barsine cardinalis˗anomala ‘species˗complex’ (Lepidoptera, Erebidae, Arctiinae). Zootaxa, 4358 (3), 441–461.
https://doi.org/10.11646/zootaxa.4358.3.3
Volynkin, A.V. & Černý, K. (2017d) On the taxonomy of the genus Barsine Walker, 1854 (Lepidoptera, Erebidae, Arctiinae). Far Eastern Entomologist, 346, 17–32.
https://doi.org/10.25221/fee.346.3
Volynkin, A.V. & Černý, K. (2018a) Description of the new genus Nebulene Volynkin & Černý, gen. nov., with a new species from Thailand and Vietnam (Lepidoptera, Erebidae, Arctiinae). Zootaxa, 4394 (2), 219–226.
https://doi.org/10.11646/zootaxa.4394.2.4
Volynkin, A.V. & Černý, K. (2018b) Revision of the Barsine zebrina species-complex, with description of three new species (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Zootaxa, 4402 (2), 339–352.
https://doi.org/10.11646/zootaxa.4402.2.6
Volynkin, A.V. & Černý, K. (2018c) Two new dark colored species of the genus Barsine Walker, 1854 from China and Indochina (Lepidoptera, Erebidae, Arctiinae, Lithosiini). Ecologica Montenegrina, 19, 61–68.
https://doi.org/10.11646/zootaxa.4586.2.8
Volynkin, A.V., Černý, K. & Ivanova, M.S. (2018) Revision of the Barsine punicea Moore, 1878 species-group, with descriptions of two new species and two new subspecies (Lepidoptera, Erebidae, Arctiinae). Journal of Asia-Pacific Entomology, 21, 999–1008.
https://doi.org/10.1016/j.aspen.2018.07.020
Volynkin, A.V., Dubatolov, V.V. & Kishida, Y. (2017) Barsura Volynkin, Dubatolov & Kishida, gen. nov., with descriptions of three new species (Lepidoptera, Erebidae, Arctiinae). Zootaxa, 4299 (1), 054–074.
https://doi.org/10.11646/zootaxa.4299.1.2
Walker, F. (1854) List of the specimens of lepidopterous insects in the collection of the British Museum. Vol. 2. Trustees of the British Museum, London, 303 pp. [pp. 279–581]
Walker, F. (1862) Catalogue of the Heterocerous Lepidopterous Insects collected at Sarawak, in Borneo, by Mr. A. R. Wallace, with Descriptions of New Species. Journal of the proceedings of the Linnean Society, Zoology, 6, 82–145.
https://doi.org/10.1111/j.1096-3642.1862.tb00935.x
Wileman, A.E. (1910) New Lepidoptera-Heterocera from Formosa. The Entomologist, 43 (567), 220–223.
Wileman, A.E. & West, R.J. (1928) New species of Heterocera from Formosa and the Philippines. The Annals and Magazine of Natural History including Zoology, Botany, and Geology, Series 10, 2, 215–224.
https://doi.org/10.1080/00222932808672870
Wu, Sh., Fu, Ch.-M., Chang, W.-Ch. & Shih, L.-Ch. (2013) Arctiinae. In: Fu, Ch.˗M., Ronkay, L. & Lin, H.˗H. (Eds.), Moths of Hehuanshan. Endemic Species Research Institute, Nantou, pp. 308–333.
Wu, S. & Kishida, Y. (2018) Elucidating the taxonomic problems of Stigmatophora karenkonis (Matsumura, 1930), comb. nov. and Miltochrista karenkensis Matsumura, 1930, stat. rev. in Taiwan (Erebidae, Arctiinae, Lithosiini). Tinea, 24 (2), 130–135.