Abstract
Key to the 22 species of genus Pheropsophus Solier 1833 of Indian subcontinent with diagnostic characters, notes, new synonyms, distribution patterns and record of two new species—Pheropsophus devagiriensis sp. nov. and P. indicus sp. nov.—from south west India are provided. Synonymisation of Pheropsophus discicollis (Dejean 1826) with Pheropsophus hilaris (Fabricius 1798) syn. nov., clearing the complications regarding the taxonomy of P. hilaris and P. sobrinus by tracking down the type series of P. hilaris, and reinstating the species status of Pheropsophus sobrinus (Dejean 1826) are the other outcomes of the present study.
Out of the 22 species recorded in India, 14 species are confined to Oriental realm and rest of the eight species penetrates into the Palearctics also; 10 (of the 22) are endemic to Indian subcontinent with P. indicus sp. nov. endemic to south western India; P. devagiriensis sp. nov. endemic to the Western Ghats; P. andrewesi, P. krichna and P. prophylax are endemic to the Indo-Burma; and P. nanodes and P. scythropus are endemic to the Eastern Himalayas.
References
Andrewes, H.E. (1919) On the types of Oriental Carabidae in the British Museum and in the Hope Department of the Oxford University Museum. Transactions of Entomological Society of London, 1919, 119–217.
https://doi.org/10.1111/j.1365-2311.1919.tb00006.x
Andrewes, H.E. (1921a) Notes on Synonymy and on some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London, 1921, 145–195.
https://doi.org/10.1111/j.1365-2311.1921.tb02805.x
Andrewes, H.E. (1921b) The Carabidae of Barkuda Island. Records of the Indian Museum, 22, 339–348.
https://doi.org/10.5962/bhl.part.1476
Andrewes, H.E. (1923a) On the Types of Carabidae described by Schmidt-Goebel in his Faunula Coleopterorum Birmaniae. Transactions of Entomological Society of London, 1923, 1–63.
https://doi.org/10.1111/j.1365-2311.1923.tb03325.x
Andrewes, H.E. (1923b) Papers on Oriental Carabidae—X. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 9 (12), 212–223.
Andrewes, H.E. (1924a) Mission Guy Babault dans les Provinces Centrales de l’Inde et dans la région occidentale de l’Himalaya, 1914. Résultats scientifiques. Insectes coléoptères Carabidae, Lahure, 1924, 1–125.
Andrewes, H.E. (1924b) On the Oriental Carabidae of the "Reise Novara". Transactions of the Entomological Society of London, 1923, 459–468.
Andrewes, H.E. (1924c) Papers on Oriental Carabidae—XIV. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 9 (14), 585–593.
Andrewes, H.E. (1927) Papers on Oriental Carabidae—XIX, The Annals and Magazine of Natural History including Zoology, Botany and Geology, 9 (19), 97–111.
Andrewes, H.E. (1929) The fauna of British India, including Ceylon and Burma. Coleoptera. Carabidae. Vol. I. Carabinae. Taylor & Francis, London, xviii + 431 pp., 9 pls.
Andrewes, H.E. (1930) Catalogue of Indian Insects. Part 18. Carabidae. Government of India Central Publication Branch, Calcutta, xxi + 389 pp.
Anichtchenko, A. (2018) Carabidae of the World. Available from: http://carabidae.org (accessed 3 March 2018)
Arrow, G.J. (1901) The Carabid genus Pheropsophus: Notes and descriptions of new species. Transactions of the Entomological Society of London, 1901, 193–206.
Arrow, G.J. (1915) Report on the Coleoptera collected by the British Ornithologists Union Expedition and the Wollaston Expedition in Dutch New Guinea by Gilbert J. Arrow, Guy A. K. Marshall, F.Z.S., C. J. Gahan, and K. G. Blair, B.Sc., F.E.S. Part 1. Transactions of Zoological Society of London, 1915, 497–508
Bates, H.W. (1886) On the Geodephagous Coleoptera collected by George Lewis in Ceylon. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 5 (17), 199–212.
https://doi.org/10.1080/00222938609460134
Bates, H.W. (1889) Contributions à la Faune Indo-Chinoise (Carabidae). Annales de la Société entomologique de France, 9 (6), 261–286.
Bates, H.W. (1891a) List of the Carabidae (Ord. Coleoptera) obtained by Père Cardon in Chota-Nagpore. Annales de la Société Entomologique de Belgique, Comptes-Rendus des Séances, 35, 324–340.
Bates, H.W. (1891b) Coleoptera In: Supplementary appendix to travels amongst the Great Andes of the Equator by Edward Whymper with contributions by H.W. Bates, T.G. Bonney, G.A. Boulenger, Peter Cameron, F. Day, W.L. Distant, A.E. Eaton, F.D. Godman, H.S. Gorham, Martin Jacoby, E.J. Miers, A. Sidney Ollif, O. Salvin, David Sharp, T.R.R. Stebbing. Illustrated. John Murray, London, pp. 7–39.
Bates, H.W. (1892a) Coléoptères du Bengale Occidental, Seconde liste des Carabidae. Annales de la Société entomologique de Belgique, 36, 230–233.
Bates, H.W. (1892b) List of Carabidae. Viaggio Di Leonardo Fea in Birmania e Regioni Vicine. Annali Del Museo Civico di Storia Naturale di Genova, 12 (2), 267–428.
Bedel, L. (1914) Catalogue raisonne des coleopteres du nord de l’Afrique. (Maroc, Algerie, Tunisie et Tripolitaine) avec notes sur la faune des iles Canaries et de Madere, premiere partie. Societe Entomologique de France, Paris, 24 pp. [pp. 297–320]
https://doi.org/10.5962/bhl.title.9132
Bouchard, J. (1903) Insectes recueillis par M. le Professeur Dr. Forster à Bornéo, Java et Sumatra (Palembang), Coléoptères Carabiques. Annales de la Société entomologique de France, 72, 169–176.
Chaudoir, M. (1843) Carabiques nouveaux. Bulletin de la Société Impériale des Naturalistes de Moscou, 16, 671–791.
Chaudoir, M. (1850) Supplément a la faune des carabiques de la Russie. Bulletin de la Société Impériale des Naturalistes de Moscou, 23 (3), 62–206.
Chaudoir, M. (1875). Genres aberrants du groupe des Cymindides. Bulletin de la Société Impériale des Naturalistes de Moscou, 49, 1–61.
Chaudoir, M. (1876) Monographie de Brachynides. Annales de la Société entomologique de Belgique, 19, 11–104.
Dejean, P.F.M.A. (1825) Species général des coléoptères de la collection de M. le Comte Dejean. Tome premier. Crevot, Paris, xxx + 463 pp.
Dejean, P.F.M.A. (1826) Species général des coléoptères de la collection de M. le Comte Dejean. Tome second. Crevot, Paris, viii + 501 pp.
Dupuis, P. (1913) H. Sauter’s Formosa-Ausbeute Carabidae. Annales de la Société entomologique de Belgique, 57, 418–425.
Dupuis, P. (1914) H. Sauter’s Formosa-Ausbeute Carabidae. Annales de la Société entomologique de Belgique, 58, 29–36.
Eisner, T., Jones, T.H., Aneshansley, D.J., Tschinkel, W.R., Silberglied, R.E. & Meinwald, J. (1977) Chemistry of defensive secretions of Bombardier Beetles (Brachinini, Metriini, Ozaenini, Paussini). Journal of Insect Physiology, 23, 1383–1386.
https://doi.org/10.1016/0022-1910(77)90162-7
Erwin, T.L. (1970) A reclassification of Bombardier beetles and a taxonomic revision of the North and Middle American species (Carabidae: Brachinida). Quaestiones Entomologicae, 6, 4–215.
Erwin, T.L. (1971) Notes and corrections to a reclassification of Bombardier beetles (Carabidae, Brachinida). Quaestiones Entomologicae, 7, 281.
Fabricius, J.C. (1775) Systema Entomologiae, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Officina Libraria Kortii, Flensburgi et Lipsiae (Flensburg and Leipzig), xxxii + 832 pp.
Fabricius, J.C. (1792) Entomologia systematica emendata et aucta. Secundum classes, ordines, genera, species adjectis synonimis, locis, observationibus, descriptionibus. Tome I. Pars I. Christ. Gottl. Proft, Hafniae, xx + 330 pp.
Fabricius, J.C. (1798) Supplementum entomologiae systematicae. Proft et Storch, Hafniae, 572 pp.
Fabricius, J.C. (1801) Systema eleutheratorum secundum ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Tomus I. Bibliopolii Academici Novi, Kiliae, xxiv + 506 pp.
Felix, R.F.F.L., Farkač, J. & Suleiman, A.S. (2012) Annotated checklist of the Carabidae (Coleoptera) of the Socotra Archipelago. In: Hájek, J. & Bezděk, J. (Eds.), Insect biodiversity of the Socotra Archipelago. Acta Entomologica Musei Nationalis Pragae, 52 (Supplementum 2), pp. 75–106.
Gestro, R. (1882) Sopra alcuni Coleotteri di Birmania raccolti dal Capitano G.B. Comotto. Annali del Museo civico di storia naturale di Genova, 18, 297–317.
Habu, A. (1984) Revised and supplementary notes on and descriptions of the Truncatipennes group of Japan (IV) (Coleoptera, Carabidae). The Entomological Review of Japan, 39, 101–173.
Heller, K.M. (1903) Uber Indo-Malayische Carabidae: Lioptera und Pheropsophus. Annales de la Société Entomologique de Belgique, 47, 241–248.
Heyne, A. & Taschenberg, O. (1908) Die Exotischen Käfer in Wort und Bild. Verlag von J.F. Schreiber, Esslingen und Munchen, 262 pp.
Hope, F.W. (1838) The Coleopterist’s Manual, Part the Second, Containing The Predaceous Land And Water Beetles Linneus And Fabricius. Henry G. Bohn, York Street, Covent Garden, London, 168 pp.
Hubenthal, W. (1911) Bemerkungen zu der Carabidengruppe Brachynini und Beschreibungen des Pheropsophus ecuadoriensis und Langenhani n.sp. (Col.). Deutsche Entomologische Zeitschrift, 1911, 547–552.
Hubenthal, W. (1914) Vier neue Arten der Carabiden–Gattung Pheropsophus Solier, nebst Bemerkungen zu bekannten Arten. Deutsche Entomologische Zeitschrift, 1914, 437–448.
Hrdlička, J. (2003) Brachininae. In: Löbl, I. & Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata—Myxophaga—Adephaga. Apollo Books, Stenstrup, pp. 212–218.
Hrdlička, J. (2017) Brachininae. In: Löbl, I. & Löbl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Volume 1. Archostemata—Myxophaga—Adephaga. Revised and Updated Edition. Brill, Leiden/Boston, 471–480.
Jedlička, A. (1963) Monographie der Truncatipennen aus Ostasien. Lebiinae–Odacanthinae–Brachyninae (Coleoptera: Carabidae). Entomologisсhe Abhandlungen und Berichte aus dem Staatliches Museum für Tierkunde in Dresden, 28 (7), 269–579. [1962–1964]
Jedlička, A. (1964) Neue Carabiden aus der Palaarctischen region (Coleoptera). Annotationes Zoologicae et Botanicae, 3, 1–6.
Kirschenhofer, E. (2010) New and Little known species of Carabidae from Middle East and Southeast Asia (Coleoptera: Carabidae: Lebiini, Brachinini). Annales Historico-Naturales Musei Nationalis Hungarici, 102, 25–64.
Lacordaire, Th. J. (1854) Histoire naturelle des insectes, Genera des Coléoptères. Tome Premier. Librairie Encyclopédique de Roret, Paris, 486 pp.
Lawrence, J.F., Hastings, A.M., Daiiwitz, M.J., Paine, T.A. & Zurcher, E.J. (1999) Beetles of the World. CSIRO Publishing, Melbourne. [CD-ROM publication]
Lewis, J. (1896) On New Species of Coleoptera from Japan and Notices of Others. The Annals and Magazine of Natural History including Zoology, Botany and Geology, 6 (17), 329–343.
https://doi.org/10.1080/00222939608680376
Linnaeus, C. von. (1771) Mantissa plantarum: Generum editionis VI. et specierum editionis II. Impensis Direct. Laurentii Salvii, Holmiæ, 588 pp.
Lorenz, W. (2005) Systematic List of Extant Ground Beetles of the World. Second Edition. Published by the author, Tutzing, 530 pp.
Lorenz, W. (2016) CarabCat: Global database of ground beetles (version Sep 2013). Species 2000. Naturalis, Leiden, ISSN 2405–8858. Available from: http://www.catalogueoflife.org/col (accessed 3 March 2018)
Macleay, W.S. (1825) Number I. of Annulosa Javanica, or an attempt to illustrate the natural affinities and analogies of the insects collected in Java by Thomas Horsfield, M.D. F.L. & G.S. and deposited by him in the museum of the honourable East-India Company. Kingsbury, Parbury, and Allen, London, xii + 50 pp., 1 pls.
Maindron, M. (1898) Description de deux espèces nouvelles de Carabiques de l’Inde orientale. Bulletin de la Société entomologique de France, 1898, 130–131.
Maindron, M. (1899) Description d’une nouvelle espece de coleoptere carabique recueillie dans le Sind (Inde occidentale). Bulletin de la Societe Entomologique de France, 1899, 16–17.
Maindron, M. (1905) Notes synonymiques sur quelques coleopteres de la famille des Carabidae. Bulletin de la Societe Entomologique de France, 1905, 94–95.
Maindron, M. (1906) Description d’une nouvelle espèce de Pheropsophus et remarques sur ce genre. Bulletin de la Société entomologique de France, 1906, 15–16.
Mani, M.S. (1955) Entomological survey of Himalaya (Part IV). Agra University Journal of Research, 4 (1), 157–170.
Motschulsky, V. von (1855) Sur les collections Coléoptèrologiques de Linné et de Fabricius. Etudes Entomologiques, 4, 25–71.
Olivier, G.A. (1795) Entomologie ou Histoire naturelle des insectes, avec leurs caractères génériques et spécifiques, leur description, leur synonymie et leur figure enluminée. Lanneau, Paris, 557 pp.
Prasad, K. (1997) Distribution of Carabid Beetles (Coleoptera: Carabidae) in Karnataka with Notes on Their Habitats and Feeding Habits. Karnataka Journal of Agricultural Sciences, 10 (4), 991–998.
Putzeys, J.A.A.H. (1875) Notes sur les Carabiques recueillis par M. Jean Van Volxem à Ceylan, à Manille, en Chine et au Japon (1873–1874). Annales de la Société Entomologique de Belgique, 18, 45–55.
Redtenbacher, L. (1867) Zoologischer Theil. Zweiter Band. I. Abtheilung A.I. Coleoptera. Reise der Österreichischen Fregatte Novara un die Erde in den Jahren 1857, 1858, 1859 unter der befehlen des Commodere B. von Wüllerstorf-Urbair. Karl Gerold’s Sohn, Wien, 1–249. [includes 1844]
Saha, S.K., Mukherjee, A.K. & Sengupta, T. (1992) Records of the Zoological Survey of India. Occasional paper No. 144. Carabidae (Coleotera: Insecta) of Calcutta. A.K. Chatterjee at Jnanodaya Press, Calcutta, 63 pp.
Saha, S.K. & Halder, S.K. (2000) Insecta: Coleoptera: Adephaga: Carabidae. In: Alfred J.R.B. (Ed.), State Fauna Series 4. Fauna of Meghalaya. Part–5. Insecta. Director, Zoological Survey of India, Calcutta, pp. 1–40.
Schaum, H.R. (1847) Bemerkungen über Fabricische Käfer. Stettiner Entomologische Zeitung, 8, 39–57.
Schmidt-Göbel, H.M. (1846) Faunula Coleopterorum Birmaniae, adjectis nonnullis Bengaliae indigenis. Gottlieb Haase Söhne, Prague, viii + 94 pp., 3 pls.
Solier, M. (1833) Sur les Deux Genres Brachinus et Aptinus du Species de M. Le Comte Dejean; et Description D’une Nouvelle Espèce de Gyrinus, par M. Solier (de Marseille). Annales de la Société entomologique de France, 2, 459–465.
Ullah, M., Naeem, M., Mahmood, K. & Rafi, M.A. (2017) Faunistic studies of the tribe Brachinini (Carabidae: Coleoptera) from northern Pakistan. Zootaxa, 4232 (2), 173–184.
https://doi.org/10.11646/zootaxa.4232.2.2
Weber, F. (1801) Observationes entomologicae, continentes novorum quae condidit generum characteres, et nuper detectarum specierum descriptiones. Impensis Bibliopolii Academici Novi, Kiliae, xii + 116 pp. [DP: March 1801 (Evenhuis 1997b: 809)]
https://doi.org/10.5962/bhl.title.8639