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Type: Monograph
Published: 2009-05-11
Page range: 1–76
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Revision of the genus Harmothoe Kinberg, 1856 (Polychaeta: Polynoidae) in the Northeast Atlantic

Senckenberg, Forschungsinstitut und Naturmuseum Frankfurt, Sektion Marine Evertebraten II, Senckenberganlage 25, D-60325 Frankfurt, Germany
Senckenberg, Forschungsinstitut und Naturmuseum Frankfurt, Sektion Marine Evertebraten II, Senckenberganlage 25, D-60325 Frankfurt, Germany
Annelida Taxonomy new species neotypes identification key

Abstract

The type material and original descriptions of 126 nominal species from the Northeast Atlantic originally described as belonging to Harmothoe Kinberg, 1856 or later referred to this genus were examined. Of these species, only 28 are considered to be valid, i.e. less than 25 %, the others being either junior synonyms, indeterminable, invalid, or referred to other genera. The valid species are described and figured, including two new species, H. fernandi n. sp. and H. mariannae n. sp. For H. extenuata (Grube, 1840), H. imbricata (Linnaeus, 1767), and H. impar (Johnston, 1839), neotypes are designated, since their type material is lost and their original descriptions are insufficient. Harmothoe aequespina (Langerhans, 1884) and H. clavigera (M. Sars, 1863) represent new combinations. An illustrated identification key to all valid Harmothoe species of the region is given.

References

  1. Amoureux, L., Rullier, F. & Fishelson, L. (1978) Systématique et écologie d'annélides polychètes de la presqu'île du Sinai. Israel Journal of Zoology, 27, 57–163.

    Augener, H. (1918) Polychaeta. Beiträge zur Kenntnis der Meeresfauna Westafrikas, 2(2), 67–625.

    Augener, H. (1931) Die bodensässigen Polychäten nebst einer Hirudinee der Meteor-Fahrt. Mitteilungen aus dem Zoologischen Staatsinstitut und Zoologischen Museum in Hamburg, 44, 279–313.

    Averincev, V.G. (1977) Biocoenoses of the shelf of Franz Josef Land and the fauna of adjacent waters, About the systematic position, ecology and distribution of Lagisca extenuata (Grube, 1840), L. rarispina (Sars, 1860) and L. propinqua Malmgren, 1867 (Polychaeta, Polynoidae). Explorations of the Fauna of the Seas, 14(12), 185–193.

    Baird, W. (1865) Contributions towards a monograph of the species of annelides belonging to the Aproditacea, containing a list of the known species, and a description of some new species contained in the National Collection of the British Museum. Journal of the Linnean Society London, ser. Zool, 8, 172–202.

    Barnich, R. & Fiege, D. (2000) Revision of the Mediterranean species of Harmothoe Kinberg, 1856 and Lagisca Malmgren, 1865 (Polychaeta, Polynoidae, Polynoinae) with descriptions of a new genus and a new species. Journal of Natural History, 34, 1889–1938.

    Barnich, R. & Fiege, D. (2003) The Aphroditoidea (Annelida, Polychaeta) of the Mediterranean Sea. Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 559, 1–167.

    Barnich, R., Fiege, D. & Sun, R. (2004) Polychaeta of Hainan Island, South China Sea. Part III. Aphroditoidea (Annelida: Polychaeta). Species Diversity, 9, 285–329.

    Barnich, R., Fiege, D., Micaletto, G., & Gambi, M.C. (2006) Redescription of Harmothoe spinosa Kinberg, 1856 (Polychaeta, Polynoidae) and related species from Subantarctic and Antarctic waters, with the erection of a new genus. Journal of Natural History, 40(1–2), 33–75.

    Bidenkap, O. (1894a) Diagnoser af tre nye Annulata Polychaeta. Christiania Videnskabs-Selskabs Forhandlinger, 3, 3–6.

    Bidenkap, O. (1894b) Systematisk oversigt over Norges Annulata Polychaeta. Christiania Videnskabs-Selskabs Forhandlinger, 10, 1–142.

    Bobretzky, N. (1882) On the annelids of the Black Sea. Zapiski Kievskago Obscestva Estestvoispytatelej, 6, 183–212.

    Brito, M.C., Núñes, J. & Bacallado, J.J. (1991) Polynoidae (Polychaeta) from the Canary Islands. Bulletin Marine Sciences, 48(2), 180–188.

    Chamberlin, R.V. (1919) The Annelida Polychaeta. Memoirs of the Museum of Comparative Zoölogy, Harvard College, 48, 1–514.

    Chambers, S.J. (1989) Leucia nivea, A polynoid (Polychaeta) new to the British Isles. Irish Natural History Journal, 23(4), 145–147.

    Chambers, S.J. & Muir, A.I. (1997) Polychaetes: British Chrysopetaloidea, Pisionoidea, and Aphroditoidea. Synopses of the British Fauna, 54, 1–202.

    Claparède, E. (1868) Les Annélides chétopodes du Golfe de Naples. Mémoires de la Société de Physique et d'Histoire Naturelle de Genève, 19(2), 313–584.

    Claparède, E. (1870) Les Annélides chétopodes du Golfe de Naples. Supplement. Mémoires de la Société de Physique et d'Histoire Naturelle de Genève, 20(2), 365–542.

    Czerniavsky, V. (1882) Materialia ad zoographiam ponticam comparatam. Fasc. III. Vermes. Bulletin de la Société impériale des Naturalistes de Moscou, 57(1), 146–198.

    Dalyell, J.G. (1853) The powers of the creator displayed in the creation. Observations on the life amidst the various forms of the humbler tribes of animated nature. With practical comments and illustrations. Volume II. John van Voorst, London, 326 pp.

    Day, J.H. (1950) On the Polychaeta collected by Mr. J. Colman at St. Helena. Journal of the Linnaean Society London, Zoology, 41(1949/1950), 434–451.

    Day, J.H. (1960) The polychaete fauna of South Africa. Part 5. Errant species dredged off Cape coasts. Annals of the South African Museum, 45, 261–373.

    Day, J.H. (1963) The Polychaete fauna of South Africa. Part 8. New species from grab samples and dredgings. Bulletin of the British Museum (Natural History), London, 10(7), 390–391.

    Day, J.H. (1967) A Monograph on the Polychaeta of Southern Africa. Part I, Errantia. British Museum (Natural History), London, 458 pp.

    Delle Chiaje, S. (1830) Figure. Memorie sulla Storia e Notomia degli Animali senza Vertebre del Regno di Napoli, plates 70–109 [imprinted date 1822; plates 1–69 published in 1822 and plates 70–109 published in 1830, cf. Sherborn (1922)].

    Ditlevsen, H. (1911) Annelids from the Danmark expedition. Danmark-Ekspeditionen til Grønlands nordøstkyst 19061908, 5(9), 411–432.

    Ditlevsen, H. (1917) Annelids. The Danish Ingolf Expedition, 4 (4), 1–71.

    Ehlers, E. (1864) Die Borstenwürmer (Annelida Chaetopoda) nach systematischen und anatomischen Untersuchungen. 1. Band. Wilhelm Engelmann, Leipzig, 748 pp.

    Ehlers, E. (1913) Die Polychaetensammlungen der Deutschen Südpolar-Expedition, 1901–1903. Deutsche Südpolar-Expedition 19011903, Zoologie 5. Band, 13(4), 397–598.

    Eliason, A. (1962) Die Polychaeten der Skagerak-Expedition 1933. Zoologiska Bidrag från Uppsala, 33, 207–293.

    Fauvel, P. (1913) Quatrième note préliminaire sur les polychètes provenant des campagnes de l'"Hirondelle" et de la "Princesse-Alice", ou déposées dans le Musée Océanographique de Monaco. Bulletin de l’Institut Océanographique, Monaco, 270, 1–80.

    Fauvel, P. (1914a) Annélides Polychètes non pélagiques provenant des Campagnes de l’"Hirondelle" et de la "Princesse-Alice" (1885–1910). Résultats des campagnes scientifiques accomplies sur son yacht par Albert Ier Prince Souverain de Monaco, 46, 1–432.

    Fauvel, P. (1914b) Aphroditiens pélagiques des Campagnes de l’"Hirondelle", de la "Princesse-Alice" et de l’"Hirondelle II". (Note préliminaire). Bulletin de l’Institut Océanograpique, 287, 1–8.

    Fauvel, P. (1923) Faune de France. 5. Polychètes errantes. P. Lechevalier, Paris, 488 pp.

    Fiege, D. & Barnich, R. (in press) Polynoidae (Annelida, Polychaeta) associated with cold water coral-reefs of the Northeast Atlantic and the Mediterranean Sea. Zoosymposia.

    Giard, A. (1886) Sur quelques Polynoidiens. Bulletin Scientifique du Département du Nord et du Pays Voisin, série 2, 9, 1–18.

    Grube, E. (1840) Actinien, Echinodermen und Würmer des Adriatischen- und Mittelmeers nach eigenen Sammlungen beschrieben. J.H. Bon, Königsberg, 92 pp.

    Grube, E. (1858) Annulata Örstediana. Videnskabelige Meddelelser fra den naturhistoriske Forening i Kjöbenhavn, 1856, 44–62 [date imprinted 1856–1857, but published in 1858, cf. Wolff & Petersen (1991)].

    Grube, E. (1860) Beschreibung neuer oder wenig bekannter Anneliden. Archiv für Naturgeschichte, 26 (1), 71–118.

    Grube, E. (1863) Beschreibung neuer oder wenig bekannter Anneliden. Archiv für Naturgeschichte, 29 (1), 37–69.

    Grube, E. (1869) Beschreibungen neuer oder wenig bekannter von Hrn. Ehrenberg gesammelter Anneliden des rothen Meeres. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin, 1869, 484–521 [imprinted date 1870, but published in 1869, cf. Zoological Record (1870)].

    Hansen, G.A. (1879) Annelider fra den norske Nordhavexpedition i 1876. Nyt Magazin for Naturvidenskaberne, Oslo, 24(1), 1–14.

    Hansen, G.A. (1880) Annelider fra den norske Nordhavsexpedition i 1878. Nyt Magazin for Naturvidenskaberne, Oslo, 25, 224–234.

    Hansen, G.A. (1882) Annelida. The Norwegian North-Atlantic Expedition 18761878. Zoology, 3(2), 1–53.

    Hartman, O. (1967) Polychaetous annelids collected by the USNS Eltanin and Staten Island cruises, chiefly from Antarctic seas. Allan Hancock monographs in Marine Biology, 2, 1–387.

    Hartmann-Schröder, G. (1975) Polychaeten der Iberischen Tiefsee, gesammelt auf der 3. Reise der Meteor im Jahre 1966. Mitteilungen des Hamburger Zoologischen Museums und Institut, 72, 47–73.

    Hartmann-Schröder, G. (1996) Annelida, Borstenwürmer, Polychaeta. Die Tierwelt Deutschlands. 2. neubearbeitete Auflage, 58, 1–648.

    Haswell, W.A. (1883) A Monograph of the Australian Aphroditea. Proceedings of the Linnean Society of New South Wales, 7, 250–299.

    Holthe, T. (1977) The polychaetous annelids of Trondheimsfjorden, Norway. Gunneria, 29, 1–64.

    Hornell, J. (1891) Report on the polychaetous annelids of the L.M.B.C. district. Proceedings of the Liverpool Biological Society, 5, 223–268.

    International Comission on Zoological Nomenclature (1992) Opinion 1666. Aphrodita imbricata Linnaeus, 1767 (currently Harmothoe imbricata) and Aphrodita minuta (currently Pholoe minuta) (Annelida, Polychaeta), specific names conserved. Bulletin of Zoological Nomenclature, 49(1), 83–84.

    Intès, A. & Le Loeuff, P. (1975) Polychètes errantes de la Côte d'Ivoire. Cahiers O.R.S.T.O.M., série Océanographie, 13(4), 267–321.

    Johnston, G. (1839) Miscellanea Zoologica. VI. On the British Aphroditaceae. Annals and Magazine of Natural History, 2(12), 424–441.

    Johnston, G. (1865) A catalogue of the British non-parasitical worms in the collection of the British Museum. Trustees of the British Museum, London, 365 pp.

    Kinberg, J.G.H. (1856) Nya slägten och arter af Annelider. Öfversigt af Kongl. Vetenskaps-Akademiens Förhhandlingar Stockholm, 12(9–10), 381–388 [read 1855; printed 1856].

    Kirkegaard, J.B. (1980) Abyssal benthic polychaetes from the northeast Atlantic Ocean, southwest of the British Isles. Steenstrupia 6(8), 81–98.

    La Greca, M. (1949) Note sur les polychètes du Bosphore. Revue de la Faculté des Sciences de l’Université d’Istanbul, Série B, 14(3), 153–169.

    Lamarck, C. de (1818) Histoire naturelle des animaux sans vertèbres. 5. Deterville et Verdière, Paris, 612 pp.

    Langerhans, P. (1880) Die Wurmfauna Madeiras, II. Zeitschrift für wissenschaftliche Zoologie, 33, 271–316.

    Langerhans, P. (1884) Die Wurmfauna von Madeira, IV. Zeitschrift für wissenschaftliche Zoologie, 40(2), 247–285.

    Lankester, E.R. (1867) XIII. On some new British Polynoina. Transactions of the Linnaean Society, 25, 373–378 [read 1866, printed 1867].

    Levinsen, G.M.R. (1884) Systematisk-geografisk oversigt over de nordiske Annulata, Gephyrea, Chaetognathi og Balanoglossi. Videnskabeliga Meddelelser fra den Naturhistoriske Forening i Kjøbenhaven, 1883, 92–350.

    Linnaeus, C. (1767) Systema Naturae. 12th edition. 1(2). Holmiae Impensis Direct. Laur. Salvii, 533–1327.

    López, E. & San Martín, G. (1996) A new species of Harmothoe (Polychaeta, Polynoidae) from the Chafarinas Islands (Alboran Sea, Western Mediterranean). Cahiers de Biologie Marine, 37, 137–187.

    Loshamn, A.A. (1980) En systematisk og dyregeografisk Undersökelse over Skjellrygg-Gruppen (Familie Aphroditidae sensu Fauvel 1923). Dissertation. Zoologisk Institutt, Universitete i Oslo, 184 pp. [unpublished].

    Loshamn, A.A. (1981) Description of five polynoid species (Polychaeta) from the coasts of Norway and Sweden, including three new species, one new genus and one new generic name. Zoologica Scripta, 10, 5–13.

    Malmgren, A.J. (1866) Nordiska Hafs-Annulata. Öfversigt af Konglia Vetenskaps-Akademiens Förhhandlingar Stockholm, 22(1), 51–110 [read 1865, printed 1866].

    Malmgren, A.J. (1867) Annulata polychaeta Spetsbergiae, Groenlandiae, Islandiae et Scandinaviae hactenus cognita. Öfversigt af Kongl. Vetenskaps-Akademiens Förhhandlingar Stockholm, 24, 127–235.

    Marenzeller, E. von (1874) Zur Kenntniss der adriatischen Anneliden. Sitzungsberichte der mathematischen-naturwissenschaftlichen Classe der kaiserlichen Akademie der Wissenschaften, Wien, 69(1) (1–5), 407–482.

    Marenzeller, E. von (1876) Zur Kenntniss der adriatischen Anneliden. Zweiter Beitrag. (Polynoinen, Hesioneen, Sylliden.). Sitzungsberichte der mathematischen-naturwissenschaftlichen Classe der kaiserlichen Akademie der Wissenschaften, Wien, 72(1)(1–5), 129–171.

    Martín, D., Rosell, D. & Uriz, M.J. (1992) Harmothoe hyalonemae sp. nov. (Polychaeta, Polynoidae), an exclusive inhabitant of different Atlanto-Mediterranean species of Hyalonema (Porifera, Hexactinellida). Ophelia, 35(3), 169–185.

    McIntosh, W.C. (1874a) XXVIII. On the invertebrate fauna and fishes of St. Andrews. Annals and Magazine of Natural History, series 4, 14, 192–207.

    McIntosh, W.C. (1874b) XXXIV. On the Annelida of the Gulf of St. Lawrence, Canada. Annals and Magazine of Natural History, series 4, 13, 261–270.

    McIntosh, W.C. (1876a) On British Annelida. Part I. Euphrosynidae, Amphinomidae, Aphroditidae, Polynoidae, Acoetidae, and Sigalionidae. Transactions of the Zoological Society London, 9(7), 371–394.

    McIntosh, W.C. (1876b) On the Annelida of the "Porcupine" Expeditions of 1869 and 1870. Part I. Euphrosynidae, Amphinomidae, Aphroditidae, Polynoidae, Acoetidae, and Sigalionidae. Transactions of the Zoological Society London, 9(8), 395–416.

    McIntosh, W.C. (1885) Report on the Annelida Polychaeta collected by H.M.S. "Challenger" during the years 1873–76. Reports on the scientific results of the voyage of H.M.S. "Challenger", Zoology, 12, 1–554.

    McIntosh, W.C. (1897a) Note on Irish annelids in the Museum of Science and Art, Dublin. No. I. Scientific Proceedings, Royal Dublin Society, 8(5), 399–404.

    McIntosh, W.C. (1897b) Notes from the Gatty Marine Laboratory, St. Andrews, No. XVIII. Annals and Magazine of Natural History, series 6, 20, 167–178.

    McIntosh, W.C. (1900) Polychaeta. Amphinomidae to Sigalionidae. A Monograph of the British Annelids, 1(2), 215–444.

    McIntosh, W.C. (1919) Notes from the Gatty Marine Laboratory, St. Andrews, No. XLII. Annals and Magazine of Natural History, series 9, 3, 125–164.

    Michaelsen, W. (1896) V. Die Polychaetenfauna der deutschen Meere einschließlich der benachbarten und verbindenden Gebiete. Beiträge zur Fauna der südöstlichen und östlichen Nordsee, 2, 1–216.

    Monro, C.C.A. (1939) Part 4. Polychaeta. B.A.N.Z. Antarctic Research Expedition (19291931) Reports, series B, 4(4), 87–156.

    Moore, J.P. (1902) Description of some new Polynoidae, with a list of other Polychaeta from North Greenland waters. Proceedings of the Academy of Natural Sciences Philadelphia, 258–278.

    Moore, J.P. (1910) The polychaetous annelids dredged by the U.S.S. "Albatross" off the coast of southern California in 1904, II. Polynoidae, Aphroditidae and Segaleonidae. Proceedings of the Academy of Natural Sciences Philadelphia, 62, 328–402.

    Morgera, A. (1918) Di un nuovo Polinoide del Golfo di Napoli. Pubblicazione della Stazione Zoolgica di Napoli, 2, 1–14.

    Müller, O.F. (1771) Von Würmern des süssen und salzigen Wassers. Heineck & Faber, Kopenhagen, 200 pp.

    Müller, O.F. (1776) Zoologiae Danicae Prodomus, seu Animalium Daniae et Norvegiae indigenarum. Characteres, nomina, et synonyma imprimis popularum. Havniae, typis Hallageriis, 274 pp.

    Nolte, W. (1936) Annelidenlarven. Nordisches Plankton, 23, 59–169.

    Pallas, P.S. (1766) Miscellanea Zoologica, quibus novae imprimis atque obscurae animalium species describunture et observationibus iconibusque illustrantur. Hagae Comitum, 224 pp.

    Pennant, T. (1777) British Zoology. 4th Edition. B. White, London, 156 pp.

    Pettibone, M.H. (1953) Some scale-bearing polychaetes of Puget Sound and adjacent waters. University of Washington Press, Seattle, 89 pp.

    Pettibone, M.H. (1954) Marine polychaete worms from Point Barrow, Alaska, with additional records from the North Atlantic and North Pacific. No. 3324. Proceedings of the United States National Museum, 103, 203–356.

    Pettibone, M.H. (1963) Marine polychaete worms of the New England region. 1. Aphroditidae through Trochochaetidae. U.S. National Museum Bulletin, 227(1), 1–356.

    Pettibone, M.H. (1985) Polychaete worms from a cave in the Bahamas and from experimental wood panels in deep water of the North Atlantic (Polynoidae, Macellicephalinae, Harmothoinae). Proceedings of the Biological Society Washington, 98(1), 127–149.

    Pettibone, M.H. (1993a) Scaled polychaetes (Polynoidae) associated with ophiuroids and other invertebrates and review of species referred to Malmgrenia McIntosh and replaced by Malmgreniella Hartman, with descriptions of new taxa. Smithsonian Contributions to Zoology, 538, 1–92.

    Pettibone, M.H. (1993b) Revision of some species referrred to Antinoe, Antinoella, Antinoana, Bylgides, and Harmothoe (Polychaeta, Polynoidae, Harmothoinae). Smithsonian Contributions to Zoology, 545, 1–41.

    Potts, F.A. (1910) XII. Polychaeta of the Indian Ocean. Part II. The Palmyridae, Aphroditidae, Polynoidae, Acoetidae, and Sigalionidae. Transactions of the Linnean Society, series 2, 13, 325–353.

    Prenant, A. (1924) Andresia ampullifera nov. g., nov. sp. de la sous-famille des Polynoiniens. Bulletin de la Société Zoologique de France, 49, 19–29.

    Quatrefages, M.A. de (1866) Annélides et Géphyriens. Histoire naturelle des Annelés marins et d’eau douce, Volume 1. Librairie Encyclopédique de Roret, Paris, 588 pp. [imprinted date 1865, but published 1866, cf. Wright (1867)].

    Rathke, H. (1837) Beitrag zur Fauna der Krym. Mémoires présentés à l’Académie Impériale des Sciences de Saint-Pétersbourg, 3, 291–454.

    Reyss, D. (1961) Lagisca drachi, n. sp., polychète Aphroditidae récoltée dans le Rech Lacaze-Duthiers. Vie et Milieu, 12(3), 473–481.

    Roule, L. (1896) Annélides. Résultats scientifiques de la campagne du "Caudan" dans le Golfe de Gascogne, Août-Septembre 1895, 440–471.

    Roule, L. (1898) Notice préliminaire sur les espèces d’Annélides recueillies dans les explorations sous-marines du "Travailleur" et du "Talisman". Bulletin du Muséum d’Histoire Naturelle, 4, 190–195.

    Roule, L. (1906) Annélides et Géphyriens. Expéditions scientifiques du "Travailleur" et du "Talisman" pendant les années 1880, 1881, 1882, 1883, 1–102.

    Rullier, F. (1963) Les Annélides polychètes du Bosphore, de la Mer de Marmara et de la Mer Noire, en relation avec celles de la Méditerranée. Rapports et Procès-verbaux des Réunions de la Commission internationale scientifique pour l’exploration de la mer méditerranée, 17(2), 161–260.

    Saint-Joseph, A. de (1888) Les Annélides polychètes des côtes de Dinard. Annales des Sciences Naturelles, série 7, 5, 141–338.

    Sars, G.O. (1873) Bidrag til kundskaben om dyrelivet paa vore havbanker. Forhandlinger i Videnskabsselskabet i Kristiana, 73–119.

    Sars, M. (1861) Om de ved Norges Kyster forekommende Arter af Annelideslaegten Polynoë. Forhandlinger i Videnskabsselskabet i Kristiana, 54–62.

    Sars, M. (1863) Geologiske og zoologiske Iagttagelser, anstillede paa en Reise i en Deel af Trondhjems Stift i Sommeren 1862. II. Zoologiske Notitser fra Christiansund og Bejan. Nyt Magazin for Naturvidenskaberne, 12(3), 273–340.

    Sars, M. (1869) Fortsatte Bemaerkninger over det dyriske Livs Udbredning i Havets Dybder. Forhandlinger i Videnskabsselskabet i Kristiana, 246–275.

    Støp-Bowitz, C. (1951) Polychètes pélagiques de l’Expédition suédoise antarctique 1901–1903.‑ Further Zoologigal Results of the Swedish Antarctic Expedition 19011903, 4(7), 1–14.

    Storm, V. (1879) Bidrag til Kundskab om Throndhjemsfjordens Fauna. Det Kongelige Norske Videnskabers Selskabs Skrifter, 1878, 9–36.

    Storm, V. (1888) Direktionens Aarsberetning for 1887. Det Kongelige Norske Videnskabers Selskabs Skrifter, 1888, 77–90.

    Strøm, H. (1768) Beskrivelse over Norske Insecter. Andet Stykke. Skrifter. Kongelige Norske Videnskabers Selskabs, 4, 313–371.

    Tebble, N. & Chambers, S. (1982) Polychaetes from Scottish waters. Part 1. Family Polynoidae. Royal Scottish Museum Studies, 1–73.

    Théel, H.J. (1879) Les Annélides Polychètes des mers de la Nouvelle-Zemble. Konglia Svenska Vetenskaps-Akademiens Handlingar, Stockholm, 16(3), 1–75.

    Trautzsch, H. (1889) Beitrag zur Kenntnis der Polynoiden von Spitzbergen. Inaugural-Dissertation der philosophischen Fakultät der Universität Jena. Gustav Fischer, Jena, 44 pp.

    Uschakov, P.V. (1965) Polychaeta of the far eastern seas of the USSR. Keys to the Fauna of the USSR, 56, 1–419 [originally published 1955 in Russian, translated by the Israel Program for Scientific Translation].

    Uschakov, P.V. (1982), Polychaetes of the suborder Aphroditiformia of the Arctic Ocean and the northwestern part of the Pacific. Families Aphroditidae and Polynoidae. Fauna of the USSR. Polychaetes, 2(1), 1–271.

    Wehe, T. (2006) Revision of the scale worms (Polychaeta, Aphroditoidea) occurring in the seas surrounding the Arabian Peninsula. Part I, Polynoidae. Fauna of Arabia, 22, 23–197.

    Wesenberg-Lund, E. (1950) Polychaeta. The Danish Ingolf-Expedition, 4 (14), 1–92.