Abstract
The currently recognized robust support for the monophyly of the Lepidoptera (and the superorder Amphiesmenoptera comprising Lepidoptera + Trichoptera) is outlined, and the phylogeny of the principal lineages within the order is reviewed succinctly. The state of the taxonomic inventory of Lepidoptera is discussed separately for ‘micro-moths’, ‘macro-moths’ and butterflies, three assemblages on which work has followed historically somewhat different paths. While currently there are about 160,000 described species of Lepidoptera, the total number of extant species is estimated to be around half a million. On average, just over one thousand new species of Lepidoptera have been described annually in recent years. Allowing for the new synonyms simultaneously established, the net increase in species numbers still exceeds 800/year. Most of the additions are foreseeable in the micro-moth grade, but even for butterflies ca 100 species are added annually. Examples of particularly interesting new high-rank taxa that have been described (or whose significance has become realized) since the middle of the 20th century include the non-glossatan lineages represented by Agathiphaga and Heterobathmia and the heteroneuran families Andesianidae, Palaephatidae, Hedylidae and Micronoctuidae. Some thoughts on how present and future systematic lepidopterology might be prioritised are presented.
References
Abraham, D., Ryrholm., N., Wittzell, H., Holloway, J. D., Scoble, M. J. & Löfstedt, C. (2001) Molecular phylogeny of the subfamilies in Geometridae (Geometroidea: Lepidoptera). Molecular Phylogenetics and Evolution, 20, 65–77.
Ackery, P., de Jong, R. & vane-Wright, R. I. (1998) The butterflies: Hedyloidea, Hesperioidea and Papilionoidea. Pp. 263–300. In N.P.Kristensen (ed.) Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology, IV/35, x + 491. Walter de Gruyter, Berlin & New York.
Baldizzone, G., Landry, J.-F. & Wolf, H. van der (2006) Coleophoridae, Coleophorinae (Lepidoptera). World Catalogue of Insects 8, 1–215. Apollo Books, Stenstrup.
Beutel, R.G. & Pohl, H. (2006) Endopterygote systematics - where do we stand and what is the goal (Hexapoda, Arthropoda)? Systematic Entomology, 31, 202–219.
Braby, M.F., Villa, R. & Pierce, N.E. (2006) Molecular phylogeny and systematics of the Pieridae (Lepidoptera : Papilionoidea) : higher classification and biogeography. Zoological Journal of the Linnean Society, 147, 239–275.
Börner, C. (1925) Lepidoptera, Schmetterlinge. In: Brohmer, P.: Fauna von Deutschland (3. edn), 358–387.
Börner, C. (1939) Die Grundlagen meines Lepidopterensystems. Verhandlungen des VIII. Internationalen Kongresses für Entomologie Berlin 1938, 2, 1372–1424.
Bridges, C.A. (1988a) Catalogue of family-group and genus-group names (Lepidoptera: Rhopalocera). C. A. Bridges, Urbana, Illinois, 6 + 8 + 3 +131 + 33 + 68+ 18 + 61 + 20 + 8 pp.
Bridges, C.A. (1988b) Catalogue of Lycaenidae & Riodinidae (Lepidoptera: Rhopalocera). C. A. Bridges, Urbana, Illinois, 7 + 377 + 115 + 140 + 101 + 37 + 12 + 10 pp.
Bridges, C.A. (1988c) Catalogue of Hesperiidae (Lepidoptera: Rhopalocera). C. A. Bridges, Urbana, Illinois, vii + 205 + 67 + 78 + 54 + 20 + 7 + 8 pp.
Bridges, C.A. (1988d) Catalogue of Papilionidae & Pieridae (Lepidoptera: Rhopalocera). C. A. Bridges, Urbana, Illinois, vii + 324 + 93 + 131 + 98 + 37 + 12 + 1 + 4 + 16 pp.
Bridges, C.A. (1988e) Bibliography (Lepidoptera: Rhopalocera). C. A. Bridges, Urbana, Illinois, 392 + 139 + 45 + 4 pp.
Brock, J.P. (1971) A contribution towards an understanding of the morphology and phylogeny of the Ditrysian Lepidop-tera. Journal of Naural Hististory 5, 29–102.
Brock, J.P. (1988) Reply to Scott's Criticism. Journal of Research on the Lepidoptera, 26, 240–247.
Brock, J. P. (1990a) Pupal protrusion in some bombycoid moths (Lepidop-tera). Entomologist’s Gazette, 41, 91–98.
Brock, J.P. (1990b) Origins and phylogeny of butterflies. In: Kudrna, O. (ed.) Butterflies of Europe 2, pp. 209–233. Aula Verlag, Wies-baden.
Brown, J.W. (2005) Tortricidae (Lepidoptera). World Catalogue of Insects 5, 1–741. Apollo Books, Stenstrup.
Bucheli, S.R. & Wenzel, J (2005) Gelechioidea (Insecta: Lepidoptera) systematics: A re-examination using combined morphology and mitochondrial DNA. Molecular Phylogenetics and Evolution, 35, 380–394.
Chapman, T.A. (1917) Micropteryx entitled to ordinal rank: Order Zeugloptera. Transactions of the Entomological Society of London, 1916, 310–314.
Common, I.F.B. (1970) Lepidoptera. Pp. 765–866. In: CSIRO (sponsor): The Insects of Australia. Carlton, Melbourne University Press.
Common, I.F.B. (1990) Moths of Australia. Melbourne University Press / E. J. Brill, Leiden, NewYork, København, Köln. 535 pp, 32 pls.
Common, I.F.B. (1994) Oecophorine genera of Australia I. The Wingia group (Lepidoptera: Oecophoridae). Monographs on Australian Lepidoptera 3, i–xvi, 1–390. CSIRO Division of Entomology, Canberra.
Common, I.F.B. (1997) Oecophorine genera of Australia II. The Chezala, Philobota and Eulechria groups (Lepidoptera: Oecophoridae). Monographs on Australian Lepidoptera 5, i–xvi, 1–407. CSIRO Division of Entomology, Canberra.
Common, I.F.B. (2000) Oecophorine genera of Australia III. The Barea group and unplaced genera (Lepidoptera: Oecophoridae). Monographs on Australian Lepidoptera 8, i–xvi, 1–453. CSIRO Division of Entomology, Canberra.
Davis, D.R. (1986) A new family of monotrysian moths from austral South America (Lepidoptera: Palaephatidae), with a phylogenetic review of the Monotrysia. Smithsonian Contributions to Zoology 434, 1–202.
Davis, D.R. & Gentili, P. (2003) Andesianidae, a new family of monotrysian moths (Lepidoptera: Andesianoidea) from austral South America Invertebrate Systematics 17, 15–26
Denis, M. & Schiffermüller, I. (1775) Ankündung eines systematischen Werkes von der Schmetterlingen der Wienergegend. 323 pp., 3. Pls. Wien.
Donner, H. & Wilkinson, C. (1989) Fauna of New Zealand. Nepticulidae (Insecta: Lepidoptera). Fauna of New Zealand 16, 1–88.
Duckworth, W.D., Genoways, H.H. & Rose, C.L. (1993) Preserving natural science collections: chronicle of our environmental heritage. Washington, D.C., iii+140 pp.
Dugdale, J.S. (1974) Female genital configuration in the classification of Lepidoptera. New Zealand Journal of Zoology 1, 127–146.
Dugdale, J.S., Kristensen, N.P., Robinson, G.S. & Scoble, M.J. (1998) The smaller Microlepidoptera-grade superfamilies. Pp. 217–232. In: Kristensen, N.P. (ed.): Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Edwards, E.D., Gentili, P., Horak, M., Kristensen, N.P. & Nielsen, E.S. (1998) The Cossoid/Sesioid assemblage. Pp. 181–197. In: Kristensen, N. P. (ed.): Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Ehnbom, K. (1948) Studies on the central and sympathetic nervous system and some sense organs in the head of neuropteroid insects. Opuscula entomologica, Suppl. 8, 1–162.
Ehrlich, P.R. (1960) Foreword. Microentomology, 24 (1).
Emmet, A.M. (1992) The scientific names of British Lepidoptera: Their history and meaning. Harley Books, Colchester, 221pp.
Epstein, M.C. & J. W. Brown (2003) Early stages of the enigmatic Prodidactis mystica (Meyrick) with comments on its new family assignment (Lepidoptera: Prodidactidae). Zootaxa, 247, 1–16.
Epstein, M.C., Gertsema, H., Naumann, C. M. & Tarmann, G. M. (1998) The Zygaenoidea. Pp. 159–180. In: Kristensen, N.P. (ed.): Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Erwin, T.L. (1982) Tropical forests: their richness in Coleoptera and other arthropod species. Coleopterists’ Bulletin 36, 74–75.
Erwin, T.L . (1991) How many species are there?: revisited. Conservation Biology, 5, 1–4.
Fabricius, J.C. (1775). Systema entomologiae, sistens insectorum classes, ordines genera, species, adiectis synonymis, locis descriptionibus, observationibus. xxx+832 pp. Flensburgi et Lipsiae.
Fänger, H. (1999) Comparative morphology of tergal phragmata occuring in the dorsal thoraco-abdominal junction of ditrysian Lepidoptera (Insecta). Zoomorphology, 119, 163–183.
Fehrenbach, H. (1995) Egg shells of Lepidoptera - Fine structure and phylogenetic implications. Zoolgischer Anzeiger, 234, 19–41.
Fibiger, M. (2007) Revision of the Micronoctuidae (Lepidoptera: Noctuoidea). Part 1, Taxonomy of Pollexinae. Zootaxa, 1567, 1–116.
Fibiger, M. & Lafontaine, J.D. (2005). A review of the higher classification of the Noctuoidea (Lepidoptera) with special reference to the Holarctic fauna. Esperiana, 11, 7–92.
Fracker, S.B. (1915) The classification of lepidopterous larvae. Illinois Biological Monographsi, 2, 1–169.
Friedlander, T., Wiegmann, B. M, Regier, J.R. & Mitter, C. (1992) Phylogenetic analysis of basal Lepidoptera using multiple nuclear gene sequences. Proceedings XIX International Congress of Entomology, Beijing, China, June 28-July 4, 1992, 28.
Friedlander, T.P., Regier, J., Mitter, C.Wagner, D.L. & Fang, Q.Q. (2000). Evolution of heteroneuran Lepidoptera (Insecta) and the utility of dopa decarboxylase for Cretaceous-age phylogenetics. Zoological Journal of the Linnean Society, 130, 213–234.
Gaston, K.J. (1991a) The magnitude of global insect species richness. Conservation Biology, 5, 283–296.
Gaston, K.J. (1991b) Estimates of the near-imponderable: a reply to Erwin. Conservation Biology, 5, 564–566.
Gaston, K.J. & May, R. M. (1992) Taxonomy of taxonomists. Nature, 356, 281–282.
Gielis, C. (2003) Pterophoroidea & Alucitoidea (Lepidoptera). World Catalogue of Insects 4, 1–198. Apollo Books, Stenstrup.
Grimaldi, D. & Engel, M.S. (2005) Evolution of the insects. Cambridge University Press, Cambridge, New York etc., xv + 755 pp.
Godfray, H.C.J., Clark, B.R., Kitching, I.J., Mayo, S.J. & Scoble, M.J. (in press). The web and the structure of taxonomy. Systematic Biology.
Hammond, P. (1992) Species inventory. Pp. 17–39. In: B. Groombridge (ed.) Global Biodiversity, Status of the Earth's Living Resources. Chapman & Hall.
Hemming, F. (1937) Hübner. A bibliographical and systematic account of the entomological works of Jacob Hübner and of the supplements by Carl Geyer, Gotfried Franz von Fr¨lich and Gottlieb August Wilhelm Herrich-Schäffer.
Hennig, W. (1953) Kritische Bemerkungen zum phylogenetischen System der Insekten. Beiträge zur Entomologie, 3 (Sonderheft), 1–85.
Hennig, W. (1966) Phylogenetic Systematics. Illinois University Press, Urbana.
Heppner, J.B. (1987) Copromorphidae. Alucitidae. Carposinidae. Epermeniidae (Copromorphoidea); Glyphipterigidae. Plutellidae (Yponomeutoidea). Pp. 399–405 In F.W. Stehr (ed.) Immature Insects. Dubuque, Kendall/Hunt.
Heppner, J.B. (1991) Faunal regions and the diversity of Lepidoptera. Tropical Lepidoptera, 2 (Suppl. 1), 1–85.
Heppner, J.B. (1992) Checklist. Lepidoptera of Taiwan 1(2).
Heppner, J.B. (1995) Lacturidae, new family (Lepidoptera, Zygaenoidea). Tropical Lepidoptera, 6, 146–148.
Heppner, J.B. (1998) Classification of Lepidoptera. Part 1. Introduction. Holarctic Lepidoptera, 5 (Suppl. 1) : 1–148.
Herbulot, C. (1992) Combien d'espèces de Lépidoptères dans le monde? Alexanor, 17, 447–448.
Herrich-Schäffer, G.A.W. (1843–56) Systematische Bearbeitung der Schmetterlinge von Europa, als Text,Revision und Supplement zu J. Huebners Sammlung europäischer Schmetterlinge. Vol. 1-6. Manz, Regensburg.
Hinton, H.E. (1946) On the homology and nomenclature of the setae of lepidopterous larvae, with some notes on the phylogeny of the Lepidoptera. Transactions of the Royal Entomological Society of London, 97, 1‑37
Hinton, H.E. (1958) The phylogeny of the panorpoid orders. Annual Review of Entomology, 3, 181–206.
Hodges, R.W. (1998) The Gelechioidea. Pp. 131–158. In: Kristensen, N. P. (ed.): Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Hodges, R.W. (ed.) (1973) Check List of the Lepidoptera of America North of Mexico. E. W. Classey/Wedge Entomological Research Foundation, xxiv + 284 pp.
Hollis, D. (ed.) (1980) Animal Identification: a Reference Guide; 3, Insects. British Museum (Natural History)/John Wiley & Sons.
Holloway, J.D., Bradley, J.D. & Carter, D.J. (1987) Lepidoptera. CIE Guides to Insects of Importance to Man 1. CAB International Institute of Entomology/ Brithish Museum Natural History.
Holloway, J.D., Kibby, G. & Peggie, D. (2001) The families of Malesian moths and butterflies. Fauna Malesiana 3,i–xi, 1–455. Brill, Leiden, Boston & Köln.
Horak, M. (2006) Olethreutine moths of Australia (Lepidoptera: Tortricidae). Monographs on Australian Lepidoptera 10, i–x, 1–522. CSIRO Division of Entomology, Canberra.
Hua Li-Zhong (2005) Lepidoptera. List of Chinese Insects III. 595 pp.Sun Yat-sen University Press, Guangzhou
Hudson, G.V. (1928) The Butterflies and Moths of New Zealand. Ferguson & Osborn.
Hudson, G.V. (1938) A Supplement to the Butterflies and Moths of New Zealand. Ferguson & Osborn.
Ivanov, V.D. (1994) Comparative analysis of the wing articulation in archaic Lepidoptera. Entomologicheskoe Obozrenie 73, 569–590.
Janzen, D.H. (1988) Ecological characterization of a Costa Rican dry forest caterpillar fauna. Biotropica, 20, 120–135.
Jong, R. de, Vane-Wright, R.I. & Ackery, P. (1996) The higher classification of butterflies (Lepidoptera): problems and prospects. Entomologica scandinavica, 27, 65–101.
Kaila, L. (2004) Phylogeny of the superfamily Gelechioidea (Lepidoptera: Ditrysia): an examplar approach. Cladistics, 20, 303–340.
Karsholt, O. & Razowski, J. (eds.) 1996. The Lepidoptera of Europe. 380 pp. Apollo Books, Stenstrup.
Kitching, I.J. (1984) A historical review of the higher classification of the Noctuidae (Lepidoptera). Bulletin of the British Museum (Natural Hististory) Entomology, 49, 153–234.
Kitching, I.J. (1987) Spectacles and silver ys: a synthesis of the systematics, cladistics and biology of the Plusiinae (Lepidoptera: Noctuidae). Bulletin of the British Museum (Natural Hististory) Entomology 54, 75–261.
Kitching, I.J. & Cadiou; J.-M. (2000) Hawkmoths of the world. An annotated and illustrated revisionary checklist (Lepidoptera: Sphingidae). viii + 227 pp., 8 pls. The Natural History Museum London & Cornell University Press, Ithaca.
Kobayashi, Y. (1997) Embryogenesis of the fairy moth Nemophora albiantenella Issiki (Lepidoptera, Adelidae), with special emphasis on its phylogenetic implicatkions. International Journal of Insect Morphology and Embryology 27, 157–166
Kosztareb, M. & Schaefer, C.W. (eds) (1990) Systematics of the North American Insects and Arachnids: Status and Needs. Virginia Agricultural Experiment Station Information Series 90–1.
Kozlov, M.V., Kuznetzov, V.I. & Stekolnikov, A.A. (1998) Skeleto-muscular morphology of the pterothorax and male genitalia of Synemon plana Walker (Castniidae) and Brachodes appendiculata (Esper) (Brachodidae) with notes on phylogenetic relationships of tortricoid-grade moth families (Lepidoptera). Invertebrate Taxonomy, 12, 245–256.
Krenn, H.W. & Kristensen, N.P. (2000) Early evolution of the proboscis of Lepidoptera (Insecta): external morphology of the galea in basal glossatan moth lineages, with remarks on the origin of the pilifers. Zoologischer Anzeiger, 239, 179–196.
Krenn, H.W. & Kristensen, N.P. (2004) Evolution of proboscis musculature in Lepidoptera. European Journal of Entomology, 101, 565–575.
Kristensen, N.P. (1984) Studies on the morphology and systematics of primitive Lepidoptera (Insecta). Steenstrupia, 10, 141–191.
Kristensen, N.P. (1997) Early evolution of the Lepidoptera + Trichoptera lineage: phylogeny and the ecological scenario. Mémoires du Muséum national d’ Histoire naturelle, 173, 253–271.
Kristensen, N.P. (ed.) (1998)[1] Lepidoptera: Moths and Butterflies 1. Evolution, Systematics and Biogeography. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Kristensen, N.P. (1998) The non‑glossatan moths. Pp. 41–49. In N.P.Kristensen (ed.) Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491. Walter de Gruyter, Berlin & New York.
Kristensen, N.P. (1998) The homoneurous Glossata. Pp. 51–63. In N.P.Kristensen (ed.) Lepidoptera: Moths and Butterflies 1. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Kristensen, N.P. (1999) Phylogeny of endopterygote insects, the most successful lineage of living organisms. European Journal of Entomology 96, 237–253
Kristensen, N.P. (2003) Skeleton and muscles: adults. Pp. 39–131. In N.P.Kristensen (ed.) Lepidoptera: Moths and Butterflies 2. Handbuch der Zoologie/Handbook of Zoology IV/36, xii + 564 pp. Walter de Gruyter, Berlin & New York.
Kristensen, N.P. & Nielsen, E.S. (1979) A new subfamily of micropterigid moths from South America. A contribution to the morphology and phylogeny of the Micropterigidae, with a generic catalogue of the family (Lepidoptera: Zeugloptera). Steenstrupia, 5, 69–147.
Kristensen, N.P. & Nielsen, E.S. (1983) The Heterobathmia life history elucidated: Immature stages contradict assignment to suborder Zeugloptera (Insecta, Lepidoptera). Zeitschrift für Zoologische Systematik und Evollutionsforschung, 21, 101–124.
Kristensen, N.P. & Skalski, A.W. (1998) Phylogeny and palaeontology. Pp. 7–25. In: Kristensen, N. P. (ed.): Lepidoptera: Moths and butterflies 1. Evolution, Systematics and Biogeography. Handbuch der Zoologie/Handbook of Zoology IV/35, x + 491 pp. Walter de Gruyter, Berlin & New York.
Kuznetzov V.I. & Stekolnikov, A.A. (1979) Classification and phylogenetic relationships of the families and subfamilies of the Pyraloidea (Lepidoptera) of the Palaearctic fauna with regard of functional morphology of male genitalia. Trudy Zoologicheskogo Instituta Akademiya Nauk SSSR. Leningrad, 82, 43–73. [In Russian].
Kyrki, J. (1988) The systematic position of Wockia Heinemann, 1970, and related genera (Lepidoptera: Ditrysia: Yponomeutiae auct.). Nota Lepidopterologica, 11, 45–69.
Labandeira, C.C., Dilcher, D.L., Davis, D.R & Wagner, D.L. (1994) Ninety-seven million years of angiosperm-insect association: Paleobiological insights into the meaning of coevolution. Proceedings of the National Academy of Science, 91, 12278–12282.
Lafontaine, J.D. & Fibiger, M. (2006) Revised higher classification of the Noctuoidea (Lepidoptera). Canadian Entomologist, 138, 610–635.
Latreille, P.A. (1796) Précis des Caractères génériques des insects, dispose dans un ordre naturel par le Citoyen Latreille. 14 + 210 +5 pp. Brive, Bordeaux.
Lee, S. & Brown, R.L. (2006) A new method for preparing slide mounts of whole bodies of Microlepidoptera. journal of Asia-Pacific Entomology, 9, 249–253.
Linnaeus, C. (1758) Systema Naturae per regna tria naturae, secundum classes, ordines genera, species, cum characteribus, differentiis, synonymis, locis. 10th ed. Holmiae. 534 pp.
Löfstedt, C. & Kozlov, M.V. (2007). A phylogenetic analysis of pheromone communication in primitive moths. Pp 473–489. In: Cardé, R.T. & Minks, A.K. (eds) Insect pheromone research: new directions. Chapman & Hall, New York, xxii + 684 pp.
Löfstedt, C., Zhu, J., Kozlov, M.V., Buda, V., Jirle, E.V., Hellqvist, S., Löfqvist, J., Plass, E., Franke, S.& Francke, W. (2004) Identification of the sex pheromone of the currant shoot borer Lampronia capitella. Journal of Chemical Ecology, 30, 643–658
Lukhtanov, V.A. (2000) Sex chromatin and sex chromosome systems in nonditrysian Lepidoptera (Insecta). Journal of Zoological Systematics and Evolutionary Research, 38, 73–79.
Mayr, E., Linsley, E.G. & Usinger, R.L. (1953) Methods and Principles of Systematic Zoology. McGraw-Hill.
Mey, W. (1996) Die Köcherfliegenfauna des Fan Si Pan-Massivs in Nord-Vietnam. 1. Beschreibung neuer und endemischer Arten aus den Unterordnungen Spicipalpia und Annulipalpia (Trichoptera). Beiträge zur Entomologie, 46, 39–65
Meyrick, E. (1895) A handbook of British Lepidoptera. [viii] + 844 pp. London.
Millar J.G. (2000) Poylyene hydrocarbons and epoxides: A Second Major Class of Lepidopteran Sex Attractant Pheromones. Annual Review of Entomology, 45, 575–604.
Miller, J.S. (1991) Cladistics and classification of the Notodontidae (Lepidoptera: Noctuoidea) based on larval and adult morphology. Bulletin of the American Museum of Natural History, 204 , 1–230.
Minet, J. (1983) Etude morphologique et phylogénétique des organes tympaniques des Pyraloidea. I - généralités et homologies. (Lep. Glossata).
Minet, J. (1984) Contribution a l'analyse phylogénétique des Néolépidoptères (Lepidoptera, Glossata). Nouvelle Revue d'Entomologie (N.S.), 2, 139–149.
Minet, J. (1986) Ebauche d'une classification moderne de l'or-dre des Lépidoptères. Alexanor, 14, 291–313.
Minet, J. (1991) Tentative reconstruction of the ditrysian phylogeny (Lepidoptera: Glossata). Entomologica scandinavica, 22, 69–95.
Minet, J. (1996) Phylogeny of Ditrysia. Abstracts, XX International Congress of Entomology Florence.
Mitchell, A., Mitter, C. & Regier, J.R. (2006) Systematics and evolution of the cutworm moths (Lepidoptera: Noctuidae): evidence from two protein-coding nuclear genes´. Systematic Entomology, 31, 21–46.
Mosher, E. (1916) A classification of the Lepidoptera based on characters of the pupa. Bulletin of the Illinois State Laboratory of natural History, 12, 1–158.
Mound, L.A. (1992) Why collect? Insect Collection News, 7, 8–14.
Mound, L.A. & Gaston, K. J. (1994) Conservation and systematics – the agony and the ecstasy. Pp. 185–195 In: Gaston, K.J. , New T. R. & Samways, M. (eds) Perspectives on Insect Conservation. Intercept Andover
Naumann, C. (1971) Untersuchungen zur Systematic und Phylogenese der holarktischen Sesiidae (Insecta, Lepidoptera). Bonner zoologische Monographien 1, 1–190. [English translation (1977) Studies on the systematics and phylogeny of Holarctic Sesiidae (Insecta, Lepidoptera). xii + 208 pp., Amerind, New Delhi].
Nielsen, E.S. (1985) The monotrysiam heteroneuran phylogeny puzzle: a possible solution. Proceeding of the 3rd. Congress of European Lepidopterology, 138–143.
Nielsen, E.S., Edwards, E.D. & Rangsi, T.V. (eds) (1996) Checklist of the Lepidoptera of Australia. Monographs on Australian Lepidoptera 4, i–xiv, 1–529. CSIRO Division of Entomology, Canberra.
Nielsen, E.S. & Kristensen, N.P. (1996) The Australian moth family Lophocoronidae and the basal phylogeny of the Lepidoptera-Glossata. Invertebrate Taxonomy, 10, 1199–1302.
Nielsen, E. S & Mound, L. A. (2000). Global diversity of insects: the problems of estimating numbers. Pp. 213–222. In Raven, P. H.& Williams, T. (eds). Nature and human society: the quest for a sustainable world. Proceedings of the 1997 Forum on Biodiversity. National Academy Press, Washington, DC., i–xii, 1–625.
Ochsenheimer, F. (1807–1835). Die Schmetterlinge von Europa 1–10. (vol. 5–10, 1825–1836 by F. Treitschke). Er. Fleischer, Leipzig.
Pitkin, L.M. (2002) Neotropical ennomine moths: a review of the genera (Lepidoptera: Geometridae). Zoological Journal of the Linnean Society, 135, 121–401.
Pitkin, L.M., Han, H. & James, S. (2007) Moths of the tribe Pseudoterpnini (Geometridae: Geometrinae): a review of the genera. Zoological Journal of the Linnean Society, 150, 343–412.
Platnick, N.I. (1991) Patterns of biodiversity: tropical vs temperate. Journal of Naural Hististory, 25, 1083–1088.
Poole, R.W. (1989) Noctuidae. In: Heppner, J. B. (ed.): Lepidopterorum Catalogus (New Series) 118, 1–3, 1–1314. E. J. Brill / Flora & Fauna Publications, Leiden, New York København & Köln.
Prins, W. de & J. de Prins (2005) Gracillariidae (Lepidoptera). World Catalogue of Insects 6, 1–502. Apollo Books, Stenstrup.
Robbins, R.K. (1987) Logic and phylogeny: A critique of Scott's phylogenies to the butterflies and Macrolepidoptera. Journal of the Lepidopterists’ Society 41, 214–216.
Robinson, G.S. (1976) The preparation of slides of Lepidoptera genitalia with special reference to Microlepidoptera. Entomologis’ s Gazette, 27, 127–132.
Robinson, G.S. & Nielsen, E.S. (1993) Tineid genera of Australia. Monographs on Australian Lepidoptera, 2, 1–344.
Sauter, W. & Hättenschwiler, P. (1991) Zum System der palaearktischen Psychiden (Lep. Psychidae). 1. Teil: Liste der palaearktischen Arten. Nota lepidopterologica, 14, 69–89.
Schrank, F. (1802) Fauna Boica 2 (1–2). Nürnberg.
Scoble, M.J. (1983) A revised cladistic classification of the Nepticulidae (Lepidoptera) with descriptions of new taxa mainly from South Africa. Transvaal Museum Monograph 2, i–xi, 1–105.
Scoble, M.J. (1986), The structure and affinities of the Hedyloidea: a new concept of the butterflies. Bulletin of the British Museum (Natural History) (Entomology), 53, 251–186.
Scoble, M.J. (1992) The Lepidoptera: Form, Function and Diversity. 404 pp., Oxford University Press.
Scoble, M.J. (2003) Changing roles and perceptions in European natural history collections: from idiosynchracy to infrastructucture. In: Scoble, M. J. (ed.): ENHSIN: The European Natural History Specimen Information Network. The Natural History Museum, London, pp. 11–20.
Scoble, M.J. , Gaston, K.J. & Crook, A. (1995) Using taxonomic data to estimate species richness in Geometridae. Journal of the Lepidopterists’ Society, 49, 136–147.
Scott, J.A. (1985) The phylogeny of butterflies (Papilionoidea and Hesperioidea). Journal of Research on the Lepidoptera, 23, 241–281.
Scott, J.A. (1986) On the monophyly of the Macrolepidoptera, including a reassessment of their relationship to Cossoi-dea and Castnioidea, and a reassignment of Mimallonidae to Pyraloidea. Journal of Research on the Lepidoptera, 25, 30–38.
Scott, J.A. (1989) Parallelism and Phylogenetic Trees. Journal of Research on the Lepidoptera 27, 257–258.
Scott, J.A. & Wright, D. M. (1990) Butterfly phylogeny and fossils. In: Kudrna, O. (ed.) Butterflies of Europe 2, pp. 152–208. Aula Verlag, Wiesbaden.
Shields, O. (1993) Is Micropterigidae or Agathiphagidae the most primitive lepidopteran family? Tyô to Ga, 44, 152–156.
Stehr, F. (ed.) (1987) Immature Insects 1. Kendall/Hunt.
Strong, D.R., Lawton, J.H. & Southwood, R. (1984) Insects on Plants. Blackwell.
Vane-Wright, R.I. (2003) Evidence and identity in butterfly systematics. Pp 477–513. In Boggs, C.L., Watt, W.B. & Ehrlich, P.R. (eds) Butterflies: ecology and evolution taking flight. University of Chicago Press, Chicago.
Vane-Wright, R.I. (2007) Linnaeus’ butterflies. Pp 59–74. In Gardiner, M. & Morris, M. (eds). The Linnean Collections. The Linnean Special Issue 7, 84 pp. The Linnean Society of London.
Vane-Wright, R.I. & Ackery, P. (eds) (1989) Biology of Butterflies (2nd ed.). Academic Press.
Vári, L. (1961) South African Lepidoptera I, Lithocolletidae. Transvaal Museum Memoirs 12, i–xiv, 1–238, pls 1–112. Transvaal Museum, Pretoria.
Vári, L., Kroon, D. M. & Krüger, M. (2002) Classification and checklist of the species of Lepidoptera recorded in Southern Africa. Published by the authors, xxi + 385 pp.
Wagner, T, Neinhuis, C. & Barthlott, W. (1996) Wettability and contaminability of insect wings as function of their surface sculpures. Acta Zoologica, 77, 213–225.
Wahlberg, N. (2006) That awkward age for butterflies: Insight from the age of the butterfly subfamily Nymphalinae (Lepidoptera: Nymphalidae). Systematic Biology, 55, 703–714.
Wahlberg, N., Braby, M. F., Brower, A.V.Z., de Jong, R., Lee, M.-M., Nylin, S., Pierce, N.E., Sperling, F.A.H., Vila, R., Warren, A.D. & Zakharov, E. (2005) Synergistic effects of combining morphological and molecular data in resolving the phylogeny of butterflies and skippers. Proceedings of the Royal Society B, 272, 1577–1586.
Weller, S.J. & Pashley, D.P. (1995) In search of butterfly origins. Molecular Phylogenetics and Evolution, 4, 235–246.
Weller, S.J. , Martin, J.A. & Constable, J.L. (1994) Phylogeny of noctuoid moths and the utility of combining independent nuclear and mitochondrial genes. Systematic Biology, 43, 194–211.
Whalley, P. (1971) The Thyrididae (Lepidoptera) of Africa and its Islands. A taxonomic and zoogeographic study. Bulletin of the British Museum (Natural Hististory) Entomology. Suppl. 17, 1–198.
Wheeler, Q.D. & Meier, R. (eds) (2000) Species Concepts and Phylogenetic Theory. A Debate. Columbia University Press.
Whitfield, J.B. & Kjer, K.M. (2007 [2008]) Ancient rapid radiations of insects: challenges for phylogenetic analysis. Annual Review of Entomology, 53, 23.1–23.24 [doi 10.1146/annurev.ento.53.103106.093304]
Wiegmann, B.M., Regier, J.C. & Mitter, C. (2002) Combined molecular and morphological evidence on the phylogeny of the earliest lepidopteran lineages. Zoologica Scripta, 31, 67–81.
Zagulajev, A.K. & Sinev. S.Y. (1988) Catapterigidae fam. n. - a new family of lower Lepidoptera (Lepidoptera, Dacnonypha). Éntomologischeskoe Obozrenie 67: 593–601. [In Russian; English translation Entomological Review, 68, 35–43, 1989].