Abstract
The extant fly family Sciaridae currently comprises a great quantity of fossil species (Evenhuis, 1994; Röschmann & Mohrig, 1995; fossilworks Gateway to the Paleobiology Database). Many of them were originally attributed to the genus Sciara, the oldest one being Sciara burmitina Cockerell, 1917 from the mid-Cretaceous Burmese amber. Mohrig & Röschmann (1994) revised several Sciara species described from the Middle Eocene Baltic amber and transferred them to different extant sciarid genera. From France, only compression fossils have been described by Théobald (1937), one from the Late Eocene and seven from the Oligocene. Here we describe the first Sciaridae from the Lowermost Eocene Oise amber (France). Blagoderov et al. (2010) previously described a Lygistorrhinidae sciaroid from the same amber.
References
Blagoderov, V., Hippa, H. & Nel, A. (2010) Parisognoriste, a new genus of Lygistorrhinidae (Diptera, Sciaroidea) from the Oise amber with redescription of Palaeognoriste Meunier. ZooKeys, 50, 79–90.
https://doi.org/10.3897/zookeys.50.506
Chandler, P.J. (2002) Heterotricha Loew and allied genera (Diptera: Sciaroidea): offshoots of the stem group of Mycetophilidae and/or Sciaridae? Annales de la Société Entomologique de France, (N.S.), 38 (1–2), 101–144.
Cockerell, T.D.A. (1917) Fossil insects. [+ Appendix]. Annals of the Entomological Society of America, 10, 1–22.
https://doi.org/10.1093/aesa/10.1.1
Cumming, J.M. & Wood, D.M. (2017) Adult morphology and terminology. In: Kirk-Spriggs, A.H. & Sinclair, B.J. (Eds.), Manual of Afrotropical Diptera, Volume 1. Introductory chapters and keys to Diptera families. Suricata, Pretoria, pp. 89–134.
Evenhuis, N.L. (1994) Catalogue of the fossil flies of the world (Insecta: Diptera). Backhuys Publishers, Leiden, 600 pp.
Fossilworks Gateway to the Paleobiology Database (2019) Available from: http://fossilworks.org (Accessed 5 June 2019)
Hippa, H. & Vilkamaa, P. (2005) The genus Sciarotricha gen. n. (Sciaridae) and the phylogeny of recent and fossil Sciaroidea (Diptera). Insect Systematics & Evolution, 36, 121–144.
https://doi.org/10.1163/187631205788838492
Jaschhof, M. & Didham, R.K. (2002) Rangomaramidae fam. nov. from New Zealand and implications for the phylogeny of the Sciaroidea (Diptera: Bibionomorpha). Studia Dipterologica, Supplement, 11, 1–60.
Jaschhof, M. & Hippa, H. (2003) Sciaroid but not sciarid: a review of Ohakunea Tonnoir & Edwards, with the description of two new species (Insecta: Diptera: Bibionomorpha). Entomologische Abhandlungen, Dresden, 60, 23–44.
Jaschhof, M. & Jaschhof, C. (2008) The Ohakunea group in Tasmania, with description of Colonomyia tasmanica sp n. (Diptera: Sciaroidea). Beiträge zur Entomologie, 58 (2), 441–453.
Menzel, F. & Mohrig, W. (1997) Chapter 2.6 Family Sciaridae. In: Papp, L. & Darvas, B. (Eds.), Contribution to a manual of Palaearctic Diptera (with special reference to flies of economic importance). Nematocera and Lower Brachycera, Volume 2, Science Herald, Budapest, pp. 51–69.
Menzel, F. & Mohrig, W. (2000) Revision der paläarktischen Trauermücken (Diptera: Sciaridae). Studia Dipterologica, Supplement, Halle/Saale, 6, 1–761.
Menzel, F. & Smith, J.E. (2017) Chapter 21. Sciaridae (black fungus gnats). In: Kirk-Spriggs, A.H. & Sinclair, B.J. (Eds.), Manual of Afrotropical Diptera, Volume 2. Nematocerous Diptera and lower Brachycera. Suricata, Pretoria, pp. 557–651.
Mohrig, W. & Menzel, F. (2009) Chapter 16. Sciaridae (black fungus gnats). In: Brown, B.V., Borkent, A., Cumming, J.M., Wood, D.M., Woodley, N.E. & Zumbado, M.A. (Eds.), Manual of Central American Diptera. Volume 1. National Research Council of Canada, Monograph Publishing Program, Ottawa, pp. 279–292.
Nel, A. & Brasero, N. (2010) Oise amber. In: Penney, D. (Ed.), Biodiversity of fossils in amber from the major world deposits. Siri Scientific Press, Manchester, pp. 137–148.
Röschmann, F. & Mohrig, W. (1995) Die Trauermücken des Sächsischen Bernsteins aus dem Untermiozän von Bitterfeld/Deutschland (Diptera, Sciaridae) [The sciarid flies from Saxonian amber of Lower Miocene of Bitterfeld/Germany]. Deutsche Entomologische Zeitschrift, (N.F.), 42 (1), 17–54.
https://doi.org/10.1002/mmnd.4810420103
Shin, S., Lee, H. & Lee, S. (2019) Proposal of a new subfamily of Sciaridae (Diptera: Sciaridae), with description of one new species from South Korea. Zootaxa, 4543 (1), 127–136.
https://doi.org/10.11646/zootaxa.4543.1.8
Steffan, W.A. (1981) Chapter 15 Sciaridae. In: McAlpine, J.F., Peterson, B.V., Shewell, G.E., Teskey, H.J., Vockeroth, J.R. & Wood, D.M. (coordinators), Manual of Nearctic Diptera, Volume 1. Research Branch, Agricultural Canada Monograph 27, Ottawa, Ontario, pp. 247–255.
Théobald, N. (1937) Les insectes fossiles des terrains oligocènes de France. Bulletin Mensuel (Mémoires) de la Société des Sciences de Nancy, 1, 1–473.