Abstract
A new genus of gnaphosid spiders, Almafuerte, gen. nov., is described for seven species from Southern South America. The new genus is diagnosed by a conspicuous retrolateral tegular projection on the male palp, and by having three teeth on the cheliceral retromargin. It comprises six species from Argentina, A. peripampasica sp. nov. (type species), from Central Argentina, also present in Uruguay, A. kuru sp. nov., from Santiago del Estero, A. vigorosa sp. nov., from Corrientes, A. goloboffi sp. nov., from Salta and Jujuy, A. remota sp. nov., from Santiago del Estero and Mendoza, and A. giaii (Gerschman & Schiapelli) comb. nov., here transferred from Echemus, from Santa Fe (the female is here described by the first time); and one species from Bolivia, A. facon sp. nov., from Cochabamba. Hypotheses of relationships of the new taxa are briefly discussed, in the context of the previously proposed groupings: the subfamilies Echeminae and Drassodinae (sensu Platnick 1990) and the “Echemus group” (sensu Murphy 2007).
References
Azevedo, G.H.F., Ott, R., Griswold, C.E. & Santos, A.J. (2016) A taxonomic revision of the ground spiders of the genus Apopyllus (Araneae: Gnaphosidae). Zootaxa, 4178 (3), 301−327.
https://doi.org/10.11646/zootaxa.4178.3.1
Ferretti, N., González, A. & Pérez-Miles, F. (2012) Historical biogeography of mygalomorph spiders from the peripampasic orogenic arc based on track analysis and PAE as a panbiogeographical tool. Systematics and Biodiversity, 10 (2), 179−193.
https://doi.org/10.1080/14772000.2012.694375
Frenguelli, J. (1950) Rasgos generales de la morfología y la geología de la provincia de Buenos Aires. L.E.M.I.T. La Plata, 32, 1–72.
Gerschman de Pikelin, B.S. & Schiapelli, R.D. (1948) Arañas argentinas II. Comunicaciones del Museo Argentino de Ciencias Naturales "Bernardino Rivadavia", Zoology, 4, 1−20.
Grismado, C.J. (2008) A taxonomic revision of the spider genus Ariadna Audouin, 1826 in Argentina and Chile, with the description of five new species (Arachnida, Araneae, Segestriidae) Zoosystema, 30 (2), 333−360.
Jorge, C., López Carrión, N., Grismado, C. & Simó, M. (2013) On the taxonomy of Latonigena auricomis (Araneae, Gnaphosidae), with notes of geographical distribuion and natural history. Iheringia, Série Zoologia, 103, 66−71.
https://doi.org/10.1590/S0073-47212013000100010
Mello-Leitão, C.F. de (1943) Catálogo das aranhas do Rio Grande do Sul. Arquivos do Museu Nacional do Rio de Janeiro, 37, 147−245.
Morrone J.J. (2014) Biogeographical regionalisation of the Neotropical region. Zootaxa, 3782 (1), 1–110.
https://doi.org/10.11646/zootaxa.3782.1.1
Murphy, J. (2007) Gnaphosid genera of the world. British Arachnological Society, St Neots, Cambridgeshire 1, i−xii, 1−92; 2, i-ii, 93−605.
Ott, R. (2012) Neodrassex, a new genus of the Leptodrassex group (Araneae, Gnaphosidae) from South America. Iheringia, Série Zoologia, 102, 343−350.
https://doi.org/10.1590/S0073-47212012000300015
Ott, R. (2013) Three new species of Neodrassex (Araneae, Gnaphosidae) from Brazil. Iheringia, Série Zoologia, 103 (4), 381−387.
https://doi.org/10.1590/S0073-47212013000400008
Ott, R. & Brescovit, A.D. (2012) Arauchemus, a new spider genus of the Echemus group (Araneae: Gnaphosidae: Echeminae) from araucaria forest areas in southern Brazil, with notes on habitat preferences and phenology. Zootaxa, 3339, 44−56.
Platnick, N.I. (1975) A revision of the spider genus Eilica (Araneae, Gnaphosidae). American Museum Novitates, 2578, 1−19.
Platnick, N.I. (1983) A review of the chilensis group of the spider genus Echemoides (Araneae, Gnaphosidae). American Museum Novitates, 2760, 1−18.
Platnick, N.I. (1985) Notes on the spider genus Eilica (Araneae: Gnaphosidae). Journal of The New York Entomological Society, 93, 1073−1081.
Platnick, N.I. (1990) Spinneret morphology and the phylogeny of ground spiders (Araneae, Gnaphosoidea). American Museum Novitates, 2978, 1–42.
Platnick, N.I. & Höfer, H. (1990) Systematics and ecology of ground spiders (Araneae, Gnaphosidae) from central Amazonian inundation forests. American Museum Novitates, 2971, 1−16.
Platnick, N.I. & Murphy, J.A. (1987) Studies on Malagasy spiders, 3. The zelotine Gnaphosidae (Araneae, Gnaphosoidea), with a review of the genus Camillina. American Museum Novitates, 2874, 1−33.
Platnick, N.I. & Shadab, M.U. (1976) A revision of the Neotropical spider genus Zimiromus, with notes on Echemus (Araneae, Gnaphosidae). American Museum Novitates, 2609, 1−24.
Platnick, N.I. & Shadab, M.U. (1979) A revision of the Neotropical spider genus Echemoides, with notes on other echemines (Araneae, Gnaphosidae). American Museum Novitates, 2669, 1−22.
Platnick, N.I. & Shadab, M.U. (1981) New species and records of Neotropical Gnaphosidae (Arachnida, Araneae). Bulletin of the American Museum of Natural History, 170, 189−196.
Platnick, N.I. & Shadab, M.U. (1982) A revision of the American spiders of the genus Camillina (Araneae, Gnaphosidae). American Museum Novitates, 2748, 1−38.
Platnick, N.I. & Shadab, M.U. (1983) A revision of the Neotropical spider genus Apodrassodes (Araneae, Gnaphosidae). American Museum Novitates, 2763, 1−14.
Platnick, N.I. & Shadab, M.U. (1984) A revision of the Neotropical spiders of the new genus Apopyllus (Araneae, Gnaphosidae). American Museum Novitates, 2788, 1−9.
Ramírez, M.J. (2014) The morphology and phylogeny of dionychan spiders (Araneae: Araneomorphae). Bulletin of the American Museum of Natural History, 390, 1–374.
Ramírez, M.J. & Grismado, C.J. (2016) Description of the spider Verita williamsi, a new genus and species from Santa Fe, Argentina (Araneae, Gnaphosidae). Revista del Museo Argentino de Ciencias Naturales, Nueva Serie, 17 (2), 173−182.
Roig-Juñent, S., Flores, G., Claver, S., Debandi, G. & Marvaldi, A. (2001) Monte Desert (Argentina): insect biodiversity and natural areas. Journal of Arid Environments, 47 (1), 77−94.
https://doi.org/10.1006/jare.2000.0688
Roig-Juñent, S., Flores, G.E. & Mattoni, C. (2003) Consideraciones biogeográficas de la Precordillera (Argentina), con base en artrópodos epígeos. In: Morrone, J. & Llorente-Bousquets, J. (Eds.), Una perspectiva latinoamericana de la Biogeografía. Las Prensas de Ciencias, Facultad de Ciencias, UNAM. México, D.F., 307pp.
Vezzani, D., Schweigmann, N., Pietrokovsky, S. & Wisnivesky-Colli, C. (2001) Characterization of Triatoma guasayana Biotopes in a Hardwood Forest of Santiago del Estero, Argentina. Memorias do Instituto Oswaldo Cruz, 96 (4), 459−466.
https://doi.org/10.1590/S0074-02762001000400004
Zakharov, B.P. & Ovtsharenko, V.I. (2011) Morphological organization of the male palpal organ in Australian ground spiders of the genera Anzacia, Intruda, Zelanda, and Encoptarthria (Araneae: Gnaphosidae). Journal of Arachnology, 39, 327−336.
https://doi.org/10.1636/CA10-91.1