Open Access Open Access  Restricted Access Subscription or Fee Access

A review of the families and genera of the superfamily PLATYSCELOIDEA Bowman & Gruner, 1973 (Crustacea: Amphipoda: Hyperiidea), together with keys to the families, genera and species



The systematics and phylogenetic relationships of the families and genera of the superfamily Platysceloidea are examined, following a thorough examination of the morphology of an example of the type species of each genus, or a substitute species if the true identity of the type species is in doubt. The mouthparts are described for each type species, often for the first time, providing additional characters for phylogenetic analysis. Genera are diagnosed using the taxonomic database program DELTA (Dalwitz et al. 1999). This database is also used for a phylogenetic analysis of the genera using PAUP (Swofford 2000). Proposed taxonomic changes resulting from this study are summarised as follows. The family Pronoidae is restricted to the monotypic genus Pronoe because it has some unique characters not found in any other platysceloidean. Paralycaea, previously in Pronoidae, has characters in common with Amphithyrus and Amphithyropsis gen. nov., a new genus proposed for Paralycaea platycephala Zeidler, 1998 (here re-determined a junior synonym of Tetrathyrus pulchellus Barnard, 1930), and together they form the proposed new family Amphithyridae fam. nov. Eupronoe and Parapronoe, also previously in Pronoidae, are similar in the morphology of the mouthparts, antennae and gnathopoda, and together form the proposed new family Eupronoidae fam. nov. The family Brachyscelidae is restricted to the genus Brachyscelus because Thamneus, previously included in Brachyscelidae, has a number of characters that differ considerably from any other genus of Hyperiidea and it is therefore placed in a family of its own, Thamneidae fam. nov. The status of the family Anapronoidae, for Anapronoe, is confirmed, as is the status of the family Tryphanidae for Tryphana. The family Lycaeidae is limited to Lycaea and Simorhynchotus. The family Oxycephalidae maybe polyphyletic but more work is required to resolve the systematic status of the eight genera currently recognised. Metalycaea globosa Stephensen, 1925, sometimes included in the Oxycephalidae, is confirmed to be a junior synonym of Lycaea serrata Claus, 1879. The family Platyscelidae is restricted to four genera, Platyscelus, Paratyphis, Hemityphis and Tetrathyrus; Amphithyrus having been removed to the new family Amphithyridae. The family Parascelidae is also restricted to four genera, Parascelus, Thyropus, Schizoscelus and Euscelus. Hemiscelus, previously included in this family, is regarded a junior synonym of Hemityphis. Keys are provided for families, genera and all currently known species. All records of associations with gelatinous zooplankton are also documented, providing additional data to help resolve the phylogeny and evolutionary origins of the Hyperiidae.



Crustacea, Amphipoda, Hyperiidea, Platysceloidea, taxonomy, DELTA, phylogenetic analysis, new families, new genus, keys

Full Text:

PDF (11MB)


Barnard, K.H. (1930) Crustacea. Part X1: Amphipoda. British Antarctic (Terra Nova) Expedition 1910, Zoology, 8 (4), 307–454.

Barnard, K.H. (1931) Amphipoda. Great Barrier Reef Expedition 1928–29, Scientific Reports, 4 (4), 111–135.

Barnard, K.H. (1940) Contributions to the crustacean fauna of South Africa. XII. Further additions to the Tanaidacea, Isopoda, and Amphipoda, together with keys for the identification of the hitherto recorded marine and fresh-water species. Annals of the South African Museum, 32 (5), 381–543.

Bate, C.S. (1858) On some new Genera and Species of Crustacea amphipoda. The Annals and Magazine of Natural History, series 3, 1 (35), 361–362.

Bate, C.S. (1861) On the morphology of some Amphipoda of the Division Hyperina. Annals and Magazine of Natural History, series 3, 8 (43), 1–16, plates 1–2.

Bate, C.S. (1862) Catalogue of the specimens of Amphipodous Crustacea in the collection of the British Museum. British Museum, Natural History, London, 399 pp., plates 1–58.

Biggs, D.C. (1978) Athorybia lucida, a new species of siphonophore (Physonectae, Arthorybiidae) from the North Atlantic Ocean. Bulletin of Marine Science, 28 (3), 537–542.

Biggs, D.C. & Harbison, G.R. (1976) The siphonophore Bathyphysa sibogae Lens and Van Riemsdijk, 1908, in the Sargasso Sea, with notes on its natural history. Bulletin of Marine Science, 26 (1), 14–18.

Boeck, A. (1871) Crustacea Amphipoda Borealia et Arctica. Forhandlinger i Vedenskabs-Selskabet i Christiania Aar 1870, 81–280 (1–200).

Boeck, A. (1872/76) de Skandinaviske og Arktiske Amphipoder, i. [s.n.], Christiana. (Plates, 1872, text, 160 pp., 1876).

Bovallius, C. (1885) On some forgotten genera among the amphipodous Crustacea. Bihang till Kungliga Svenska Vetenskaps-Akademiens Handlingar, 10 (14), 1–17, 1 plate.

Bovallius, C. (1887) Systematical list of the Amphipoda Hyperiidea. Bihang till Kungliga Vetenskaps-Akademiens Handlingar, 11 (16), 1–50.

Bovallius, C. (1890) The Oxycephalids. Nova Acta Regiae Societatis Scientiarum Upsaliensis, series iii, 14, 1–141, plates 1–7.

Bowman, T.E. & Gruner, H.-E. (1973) The families and genera of Hyperiidea (Crustacea: Amphipoda). Smithsonian Contributions to Zoology, No. 146, 1–64.

Browne, W.E., Haddock, S.H.D. & Martindale, M.Q. (2007) Phylogenetic analysis of lineage relationships among hyperiid amphipods as revealed by examination of the mitochondrial gene, cytochrome oxidase 1 (CO1). Integrative and Comparative Biology, 47 (6), 815–830.

Carus, J.V. (1885) Prodromus Faunae Mediterraneae sive Descriptio Animalium Maris Mediterranei incolarum quam comparata silva rerum quatenus innotuit adjectis locis et nominibus vulgaribus eorumque auctoribus in commodum Zoologorum congessit Julius Victor Carus. Vol. I. Pars II. Arthropoda. Stuttgart, 1885.

Chevreux, E. (1892) Vibilia erratica, Amphipode pélagique nouveau du littoral des Alpes-Maritimes. Bulletin de la Société Zoologique de France, 17, 32–35, 3 Figs

Chevreux, E. (1900) Amphipodes provenant des campagnes de I'Hirondelle (1885–1888). Résultats des Campagnes scientifiques accomplies sur son Yacht, par Albert 1er, Prince Souverain de Monaco, 16 (i–iv), 1–195, plates 1–18.

Chevreux, E. & Fage, L. (1925) Amphipodes. Faune de France, 9, 488 pp., 438 text Figs

Chun, C. (1889) Bericht über eine nach den Canarischen Inseln in Winter 1887/88 ausgeführte Reise. II. Abteilung. Beobachtungen über die pelagische Tiefen und Oberflächenfauna des ostlichen atlantischen Oceans. Sitzungsberichte der Königliche Preussischen Akademie der Wissenschaften zu Berlin, 45, 519–553, plate 3.

Claus, C. (1871) Untersuchungen über den Bau und der Verwandtschaft der Hyperiden. Nachrichten von der Königliche Gesellschaft der Wissenschaften und der Georg-Augustus-Universität zu Göttingen, Yahre 1871, 149–157.

Claus, C. (1879) Die Gattungen und Arten der Platysceliden in Systematischer Übersicht. Arbeiten aus dem Zoologischen Institut der Universität zu Wien und der Zoologischen Station Triest, 2, 1–52 (147–198).

Claus, C. (1880) Grundzüge der Zoologie. 4th ed. Marburg.

Claus, C. (1884) ‘Elementary text-book of Zoology.’ General part and special part; Protozoa to Insecta.’ Translated and edited by Adam Sedgwick, M.A., with the assistance of F.G. Heathcote, B.A. London, 1884.

Claus, C. (1887) Die Platysceliden. Alfred Hölder, Vienna, 77 pp., plates 1–25.

Cocco, A. (1832) Su di alcuni nuovi Crustacei dé mari di Messina Lettera del dott. Anastasio Cocco al celebre dott. William Elford Leach uno dé conservatori del Museo britannico in Londra. Effemeridi scientifiche e letterarie per la Sicilia, 2 (6), 202–209. Giugno, 1832.

Cocco, A. (1833) Descrizione di alcuni Crustacei di Messina per Anastasio Cocco. Giornale di Scienze Lettre e Arti per la Siclia, 44 (11). Palermo, 1833.

Costa, A. (1864) Di alcuni Crostacei e di un distomideo parassito degli acalefi. Rendiconto della R. Accademia della Scienze Fisiche e Matamatiche di Napoli. Fascicolo 4.

Costa, O.G. & Costa A. (1840) Catalogo de’ Crostacei del Regno Napoli, pp. 1–7.

Dana, J.D. (1852) On the classification of the Crustacea Choristopoda or Tetradecapoda. American Journal of Sciences and Arts, series 2, 14 (41), 297–316.

Dana, J.D. (1853) Crustacea, Part II. United States Exploring Expedition, 14, 689–1618. [Plates 1–96 published in 1855]

Dallwitz, M.J., Paine, T.A. & Zurcher, E.J. (1999) User’s guide to the DELTA system. A general system for processing taxonomic descriptions. Edition 4.10. Canberra, CSIRO Division of Entomology. 156 pp.

Dick, R.I. (1970) Hyperiidea (Crustacea: Amphipoda) Keys to South African genera and species, and a distribution list. Annals of the South African Museum, 57 (3), 25–86.

Evans, F. (1967) Syntypes of Decapoda described by William Stimpson and James Dana in the collections of the British Museum (Natural History). Journal of Natural History, 1, 399–411.

Eydoux, F. & Souleyet, L.F.A. (1842) Voyage autour du monde, exécuté pendant les années 1836 et 1837 sur la corvette La Bonite, commandée par M. Vaillant. Zoologie. Tome premier, seconde partie. Arthus Bertrand, Paris, 334 pp.

Fage, L. (1960) Oxycephalidae, amphipods pélagiques. Dana Reports, 52, 1–135.

Garbowski, T. (1895) Kosmos polski. Lwowie (Lemberg), volume 21.

Gasca, R. & Haddock, S.H.D. (2004) Associations between gelatinous zooplankton and hyperiid amphipods (Crustacea: Peracarida) in the Gulf of California. Hydrobiologia, 530/531, 529–535.

Gasca, R., Hoover, R. & Haddock, S.H.D. (2014) New symbiotic associations of hyperiid amphipods (Peracarida) with gelatinous zooplankton in deep waters off California. Journal of the Marine Biological Association of the United Kingdom, 95 (3), 503–511.

Gasca, R., Suárez-Morales, E. & Haddock, S.H.D. (2006) Symbiotic associations between crustaceans and gelatinous zooplankton in deep and surface waters off California. Marine Biology (2007), 151, 233–242.

Gerstaecker, A. (1886) Dr. H.G. Bronn’s Klassen und Ordnungen des Thierreichs, wissenschaftlich dargestellt in Wort und Bilt. 5(2). Gliederfüssler; Arthropoda. Leipzig und Heidelberg, 1886, pp. 417–512.

Giles, G.M. (1888) XV.—Natural History Notes from H.M’s Indian Marine Survey Steamer ‘Investigator’, Commander ALFRED CARPENTER, R.N., Commanding. No. 6. On Six new Amphipods from the Bay of Bengal. Journal of the Asiatic Society of Bengal, 56 (2/2), 212–229, plates 3–8.

Giles, G.M. (1890) VII.—Natural History Notes from H.M’s Indian Marine Survey Steamer ‘Investigator’, Commander ALFRED CARPENTER, R.N., D.S.O. Commanding. No. 9. Further Notes on the Amphipoda of Indian waters. Journal of the Asiatic Society of Bengal, 57 (2/3), 220–255, plates 6–12.

Guérin, F.E. (1825) Encyclopédie Méthodique Histoire Naturelle. Entomologie, ou histoire naturelle des Crustacés, des Arachnides et des Insectes par M. Latreille. Tome 10. Paris.

Guérin-Méneville, F.E. (1836) Description de quelques genres nouveaux des Crustacés appartenant à la famille des Hypérines. Magasin de Zoologie, Année 6, Classe 7, 1–10, plates 17–18.

Harbison, G.R., Biggs, D.C. & Madin, L.P. (1977) The associations of Amphipoda Hyperiidea with gelatinous zooplankton – II. Associations with Cnidaria, Ctenophora and Radiolaria. Deep-Sea Research, 24, 465–488.

Harbison, G.R. & Madin, L.P. (1976) Description of the female Lycaea nasuta Claus, 1879 with an illustrated key to the species of Lycaea Dana, 1852 (Amphipoda, Hyperiidea). Bulletin of Marine Science, 26 (2), 165–171.

Harbison, G.R., Madin, L.P. & Swanberg, N.R. (1978) On the natural history and distribution of oceanic ctenophores. Deep-Sea Research, 25, 233–256.

Hope, F.W. (1851) Catalogo dei crostacei Italiani e di moltri altri del Mediterraneo. Napoli Stabilimento Tippografico di Fr’. Azzolino Vico Gerolomini, 10, 48 pp., 2 Figs

Hurley, D.E. (1955) Pelagic amphipods of the sub-order Hyperiidea in New Zealand waters. I. Systematics. Transactions of the Royal Society of New Zealand, 83 (1), 119–194.

Hurt. C., Haddock, S.H.D. & Browne, W.E. (2013) Molecular phylogenetic evidence for the reorganization of the Hyperiid amphipods, a diverse group of pelagic crustaceans. Molecular Phylogenetics and Evolution, 67, 28–37.

International Commission on Zoological Nomenclature (1950) Article 4 and Opinion 141. Conclusions of the sixth meeting of the International Commission on Zoological Nomenclature held at the Sorbonne in the Amphitheatre Louis-Liard on Thursday, 22nd July, 1948 at 1700 hours, pp. 138–139. Bulletin of Zoological Nomenclature, 4, 1–760.

International Commission on Zoological Nomenclature (1998) Opinion 1889. Parapronoe crustulum Claus, 1879 (Crustacea, Amphipoda): specific name conserved. Bulletin of Zoological Nomenclature, 55 (1), 52–53.

International Code of Zoological Nomenclature (1999) International code of Zoological Nomenclature, 4th edition. The International Trust for Zoological Nomenclature and The Natural History Museum, London, 306 pp.

Jespersen, P. & Tåning, A.V. (1934) Introduction to the Reports from the Carlsberg Foundation’s Oceanographical Expedition Round the World 1928–30 and list of stations. Dana Report, 1, 7–130.

Krumbach, T. (1911) Notizen über die Fauna der Adria bei Rovigno. IV. Die Ctenophorenfauna von Rovigno nach den Novemberstürmen 1910. Zoologischer Anzeiger, 37 (16), 315–319.

Laval, P. (1980) Hyperiid amphipods as crustacean parasitoids associated with gelatinous plankton. Oceanography and Marine Biology, Annual Review, 18, 11–56.

Lavaniegos, B.E. & Ohman, M.D. (1999) Hyperiid amphipods as indicators of climate change in the California Current, pp. 489–509. In: Schram, F.R. & Van Vaupel Klein, J.C. (Eds.). Crustaceans and the biodiversity crisis. Proceedings of the Fourth International Crustacean Congress, Amsterdam, The Netherlands, July 20–24, vol. 1. Brill, Leiden.

Lima, M.C.G. & Valentin, J.L. (2001) New records of Amphipoda Hyperiidea in association with gelatinous zooplankton. Hydrobiologia, 448, 229–235.

Lowry, J.K. (1986) The callynophore, a eucaridean/paracaridean sensory organ prevalent among the Amphipoda (Crustacea). Zoologica Scripta, 15 (4), 333–349.

Lowry, J.K. & Stoddart, H.E. (1997) Amphipoda Crustacea IV. Families Aristiidae, Cyphocarididae, Endevouridae, Lysianassidae, Scopelocheiridae, Uristidae. Memoirs of the Hourglass Cruises, 10 (1), 1–148.

Lucas. H. (1840) Histoire Naturelle des Crustacés, des Arachnides et des Myriapodes. P. Duménil, Paris, 600 pp. + 46 plates (inclusive).

Lucas, H. (1846) Les amphipodes. Histoire naturelle des animaux articulés, premiére partie Crustacés, Arachnides, Myriapodes et Hexapodes. Exploration scientifique de I'Algérie pendant les années 1840, 1841, 1842. Paris, pp. 1–88, plates 1–8.

Madin, L.P. & Harbison, G.R. (1977) The associations of Amphipoda Hyperiidea with gelatinous zooplankton – I. Associations with Salpidae. Deep-Sea Research, 24, 449–463.

Marion, A.F. (1874) Recherches sur les animaux inférieurs du golfe de Marseille. Descriptions des Crustacés Amphipodes parasites des salpes. Annales des Sciences naturelle, 6 séries Zoologie et Paléontologie, séries 5, 17, 1–19, plates 1–2.

Martin, J.W. & Davis, G.E. (2001) An updated classification of the recent Crustacea. Natural History Museum of Los Angeles County, Science Series, No. 39, 124 pp.

Milne-Edwards, H. (1830) Extrait de recherches pour servir à l'histoire naturelle des Crustacés Amphipodes. Annales des Sciences Naturelles, 20, 353–399, plates 10–11.

Milne-Edwards, H. (1838) Arachnoides, Crustacés. In: Historie Naturelle des Animaux sans Vertèbrespar. J. B. P. A. de Lamarck. Tome V. (Deuxième Edition. Par M.M.G.P. Deshayes et H. Milne-Edwards). Paris, 1838, 699 pp.

Milne-Edwards, H. (1840) Histoire naturelle des Crustacés, comprenant l’anatomie, la physiologie, et la classification de ces animaux. Tome 3. Paris, 638 pp., plates 1–42.

Montfort, P. Denys de (1810) Conchyliologie Systématique, et classification méthodique des Coquilles; offrant leurs figures, leur arrangement générique, leurs descriptions caractéristiques, leurs noms; ainsi que leur synonymie en plusieurs langues. Tome Second. Paris: F. Schoell, 1808–10, 676 pp.

Nair, K.K.C. (1993) Metalycaea globosa Stephensen, a valid species of Oxycephalidae (Amphipoda, Hyperiidea). Journal of Plankton Research, 15 (10), 1171–1176.

Nair, K.K.C. (1995) Taxonomic features and identification of Oxycephalidae (Platysceloidea, Physocephalata, Hyperiidea, Amphipoda). Mahasagar, 28 (1–2), 1–65.

Natale, G.De. (1850a) Su pochi Crostacei del porto di Messina. Lettera del Dottor Giuseppe de Natale socio corrispondente dell’ Academia Gioenia di Catania al Sig. Achille Costa con una tavola in rame. Napoli, pp. 1–16.

Natale, G.De. (1850b) Descrizione zoologica d'una nuova specie di Plojaria e di alcuni Crostacei del porto di Messina con poche considerazioni generali sulla natura delle appendici aculeiformi delle piante e degli animali. Messina, 31 pp, appendix 2 pp., plates 1–2.

Nicolet, H. (1849) Crustacéos, pp. 115–318. In: Historia fisica y politica de Chile segun documentos adquiridos en esta republica durante doce anos de residencia en ella y publicado bajo los auspicious del supremo gobierno por Claudio Gay ciudadano chileno. Zoologia, 3, Paris and Santiago, 547 pp.

Oshel, P.E. & Steele, D.H. (1988) Comparative morphology of amphipod setae, and a proposed classification of setal types. Crustaceana Supplement, 13, 100–106.

Pillai, N.K. (1966) Pelagic amphipods in the collections of the Central Marine Fisheries Research Institute, India: Part I. Family Oxycephalidae. Proceedings of the Symposium on Crustacea held at Ernakulam from January 12–15, 1965. Part I, 169–204. (Marine Biological Association of India).

Pirlot, J.M. (1929) Résultats zoologiques de la croisière atlantique de’l’Armauer Hansen’ (Mai-Juin 1922). 1. Les Amphipodes Hypérides. Mémoires de la Société Royale des Sciences de Liège, série 3, 15 (2), 1–196.

Pirlot, J.M. (1930) Les Amphipodes de l’expedition du ‘Siboga’, Première Partie, Les Amphipodes Hypérides (à l’exception des Thaumatopsidae et des Oxycephalidae). Siboga-Expeditie, Monograph, 33 (a), 1–54.

Pirlot, J.M. (1939) Sur des Amphipodes Hypérides provenant des croisières du Prince Albert 1er de Monaco. Résultats des Campagnes Scientifiques accomplies sur son Yacht par Albert 1er Prince Souverain de Monaco, fascicule 102, 1–64.

Prestandrea, N. (1833) Su di alcuni nuovi crostacei dei mari di Messina. Effemeridi Scientifiche e Letterarie per la Sicilia, 6, 3–14. Palermo.

Risso, A. (1816) Histoire naturelle des Crustacés des environs de Nice. Librairie Grecque-Latine-Allemande: Paris, 175 pp., 3 plates.

Sars, G.O. (1890) Amphipoda. In: An account of the Crustacea of Norway with short descriptions and figures of all the species. Volume 1: viii + 711 pp., 240 plates, 8 supplementary plates. Alb. Cammermeyer, Kristiana.

Schellenberg, A. (1927) Amphipoda des Nordischen Plankton, pp. 589–722. In: Nordisches Plankton, Zoologischer Teil. Kiel, Leipzig. Volume 3, 722 pp.

Semenova, T.N. (1981) Parapronoe elongata sp. n. (Crustacea, Hyperiidea) and discussion of the status of the genus Sympronoe Stebbing, 1888. Zoologicheskii Zhurnal, 10, 1581–1585. [In Russian]

Shih, C.-T. & Chen, Q.-C. (1995) Zooplankton of China Seas (2) – The Hyperiidea (Crustacea: Amphipoda). China Ocean Press, Beijing, 295 pp.

Spamer, E.E. & Bogan, A.E. (1992) General Invertebrates Collection of the Academy of Natural Sciences of Philadelphia. Tryonia, 26, 1–305.

Spamer, E.E. & Bogan, A.E. (1994) Type specimens of Crustacea surviving in the Guérin-Méneville collection, Academy of Natural Sciences of Philadelphia. Proceedings of the Academy of Natural Sciences of Philadelphia, 145, 35–46.

Spandl, H. (1924a) Expeditionen S.M. Schiff ‘Pola’ in das Rote Meer. Nördliche und Südliche Hälfte, 1895/96–1897/98. Zoologische Ergebnisse 35. Die Amphipoden des Roten Meeres. Denckschriften Akademie der Wissenschaften in Wien, Mathematisch-Naturwissenschaftliche Klasse, 99, 19–73.

Spandl, H. (1924b) Amphipoda Hyperiidea aus der Adria. Zoologischer Anzeiger, 58, 261–272.

Spandl, H. (1927) Die Hyperiiden (exkl. Hyperiidea Gammaroidea und Phronimidae) der Deutschen Südpolar-Expedition 1901–1903. Deutsche Südpolar-Expedition 1901–1903, Band 19, Zoologie, 11, 145–287, plate 10.

Stebbing, T.R.R. (1888) Report on the Amphipoda collected by H.M.S. ‘Challenger’ during the years 1873–1876. Report on the Scientific Results of the Voyage of H.M.S. ‘Challenger’ during the years 1873–76. Zoology, 29, i–xxiv & 1–1737, plates 1–210.

Stebbing, T.R.R. (1895) Descriptions of nine new species of amphipodous crustaceans from the tropical Atlantic. Transactions of the Zoological Society of London, 13 (10), 249–371, plates 51–55.

Stephensen, K. (1923) Crustacea Malacostraca, V. (Amphipoda. 1) Danish Ingolf-Expedition, 3 (8), 1–100, +2 tables, 1 map.

Stephensen, K. (1925) Hyperiidea-Amphipoda (Part 3: Lycaeopsidae, Pronoidae, Lycaeidae, Brachyscelidae, Oxycephalidae, Parascelidae, Platyscelidae). Report on the Danish Oceanographical Expeditions 1908–10 to the Mediterranean and Adjacent Seas, 2 (Biology–D5), 151–252.

Stephensen, K. (1932) The Tanaidacea and Amphipoda of the Arctic. Fauna Arctica, 6, 343–378.

Steuer, A. (1911a) Ein Vertreter der Hyperiidea curvicornia aus der Adria. Zoologischer Anzeiger, 37 (17), 351–352.

Steuer, A. (1911b) Adriatische Planktonamphipoden. Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, Wien, 120 (6), 671–688, plates 1–3.

Stewart, D.A. (1913) A report on the extra-Antarctic Amphipoda Hyperiidea collected by the ‘Discovery’. Annals and Magazine of Natural History, 8 (12), 245–264, plates 4–7.

Streets, T.H. (1877) Contributions to the Natural History of the Hawaiian and Fanning Islands and Lower California. Bulletin of the United States National Museum, 7, 1–172.

Streets, T.H. (1879) Pelagic Amphipoda. Proceedings of the Academy of Natural Sciences of Philadelphia, 1878, 276–290, plate 2.

Sudara, S. (1975) A new species of Cranocephalus (Amphipoda, Hyperiidea, Oxycephalidae) from the Gulf of Thailand and the South China Sea. Natural History Bulletin of the Siam Society, 26, 41–50.

Swofford, D.L. (2000) PAUP. Phylogenetic Analysis using parsimony (and other methods) version 4.0b4a. Sinauer Associates Inc., Sunderland, Massachusetts.

Tashiro, J.E. (1978) Comparison of Eupronoe armata Claus, 1879, and Eupronoe intermedia Stebbing, 1888 (Amphipoda, Hyperiidea). Crustaceana, 34 (1), 76–82.

Thomson, G.M. (1879) New Zealand Crustacea. Tribe III-Amphipoda. Transactions and Proceedings of the New Zealand Institute, 11, 235–248, plate 10.

Thurston, M.H. (1976) The vertical distribution and diurnal migration of the Crustacea Amphipoda collected during the SOND Cruise, 1965. II. The Hyperiidea and general discussion. Journal of the Marine Biological Association of the United Kingdom, 56, 383–470.

Vinogradov, G.M. (1990) Pelagic amphipods (Amphipoda, Crustacea) from the south-eastern Pacific. Transactions of the P.P. Shirshov Institute of Oceanology, 124, 27–104. [In Russian].

Vinogradov, G.M. (1999) Amphipoda, pp. 1141–1240. In: Boltovskoy, D. (Ed.), South Atlantic Zooplankton. Vol. 2. Backhuys, Leiden, The Netherlands, 1705 pp.

Vinogradov, M.E., Volkov, A.F. & Semenova, T.N. (1982) Amfipody-Giperiidy (Amphipoda: Hyperiidea) Mirovogo Okeanea. Akademiya Nauk SSSR, Opredeliteli po Faune SSSR No. 132. Leningrad, 492 pp. [In Russian, English translation, 1996, Smithsonian Institution Libraries, Washington D.C., D. Siegel-Causey, Scientific Editor].

White, A. (1847) List of the specimens of Crustacea in the Collections of the British Museum, Printed by the order of the Trustees, London (E. Newman), 1847, 143 pp.

Yoo, K.I. (1971) Pelagic hyperiids (Amphipoda: Hyperiidea) of the western North Pacific Ocean. Journal of the National Academy of Sciences, Republic of Korea, Natural Science Series, 10, 39–89.

Young, J.W. & Anderson, D.T. (1987) Hyperiid amphipods (Crustacea: Peracarida) from a warm-core eddy in the Tasman Sea. Australian Journal of Marine and Freshwater Research, 38, 711–725.

Zeidler, W. (1978) Hyperiidea (Crustacea: Amphipoda) from Queensland waters. Australian Journal of Zoology, Supplementary Series, No. 59, 1–93.

Zeidler, W. (1992) Hyperiid amphipods (Crustacea: Amphipoda: Hyperiidea) collected recently from eastern Australian waters. Records of the Australian Museum, 44 (1), 85–133.

Zeidler, W. (1996a) Case 2992. Parapronoe crustulum Claus, 1879 (Crustacea: Amphipoda): proposed conservation of the specific name. The Bulletin of Zoological Nomenclature, 53 (3), 178–181.

Zeidler, W. (1996b) On the identity of Typhis rapax Milne-Edwards, 1830 (Crustacea: Amphipoda: Hyperiidea). Crustaceana, 69 (6), 734–741.

Zeidler, W. (1997a) The hyperiid amphipod specimens in the Guérin-Méneville collection surviving in the collections of the Academy of Natural Sciences of Philadelphia (Crustacea: Amphipoda: Hyperiidea). Proceedings of the Academy of Natural Sciences of Philadelphia, 148, 121–146.

Zeidler, W. (1997b) A new species of Anapronoe with a redescription of A. reinhardti Stephensen, 1925 (Amphipoda: Hyperiidea: Anapronoidae). Ophelia, 47 (2), 133–150.

Zeidler, W. (1998) Pelagic amphipods (Crustacea: Amphipoda: Hyperiidea) collected from eastern and south-eastern Australian waters by the C.S.I.R.O. research vessel ‘Warreen’ during the years 1938–41. Records of the South Australian Museum. Monograph Series, No. 4, 1–143.

Zeidler, W. (1999) Review of the hyperiidean amphipod genus Oxycephalus Milne-Edwards (Crustacea: Amphipoda: Oxycephalidae). Invertebrate Taxonomy, 13, 391–424.

Zeidler, W. & Browne, W.E. (2015) A new Glossocephalus (Crustacea: Amphipoda: Hyperiidea: Oxycephalidae) from deep-water in the Monterey Bay region, California, USA, with an overview of the genus. Zootaxa, 4027 (3), 408–424.



  • There are currently no refbacks.

ISSN 1175-5326 (Print Edition) & ISSN 1175-5334 (Online Edition)
Published by Magnolia Press, Auckland, New Zealand