Skip to main content Skip to main navigation menu Skip to site footer
Type: Article
Published: 2016-11-10
Page range: 516–542
Abstract views: 397
PDF downloaded: 16

A historical review of the classification of Erebinae (Lepidoptera: Erebidae)

Entomology and Nematology Department, University of Florida, Gainesville, Florida, 32611 USA Florida Museum of Natural History, Gainesville, Florida, USA 32611
Florida Museum of Natural History, Gainesville, Florida, USA 32611
Entomology and Nematology Department, University of Florida, Gainesville, Florida, 32611 USA Florida Museum of Natural History, Gainesville, Florida, USA 32611
Lepidoptera Catocalinae Erebidae Erebinae Noctuidae Noctuoidea systematics taxonomy

Abstract

Erebidae is one of the most diverse families within the order Lepidoptera (butterflies and moths), with nearly 25,000 described species. The nominal subfamily Erebinae is among the most species rich and taxonomically complex. It reaches its highest diversity in the tropics, where much of the fauna remains undescribed. Species in this subfamily feed on a broad range of host plants, with associated radiations on grasses and legumes, and some species are pests of agriculture and forestry. The Erebinae, as defined today, comprises a large portion of the former noctuid subfamily Catocalinae. However, many lineages have tenuous or uncertain systematic placement. Here, we review the complex historical classification of Erebinae, and discuss the possible placement of some of these lineages in light of traditional morphological groupings and recent molecular phylogenetic analyses. We present an updated list of named erebine tribes and their relationships, and identify morphological traits from literature used to group genera within these tribes.

 

References

  1. Agenjo, R. (1959) Las Catocala Schrk., 1802, españolas, con más amplias consideraciones respeto a las de mayor interés forestal. Eos, 35, 301–384.

    Angulo, A.O., Jana-Saenz, C. & Parra, L.E. (1986) Estudio del metafurcasterno en algunas especies de noctuidos: valor morphofuncional (Lepidoptera: Noctuidae). Annales de la Société Entomologique de France, 22, 369–374.

    Barnes, W. & McDunnough, J.H. (1917) Check list of the Lepidoptera of Boreal America. Herald Press, Decatur, Illinois, 828 pp.
    http://dx.doi.org/10.5962/bhl.title.10097

    Beck, H. (1960) Die Larvalsystematik der Eulen (Noctuidae). Abhandlungen zur Larvalsystematik der Insekten, Vol. 4, 406pp.

    Beck, H. (1999) Die Larven der Europäischen Noctuidae. Revision der Systematik der Noctuidae (Lepidoptera: Noctuidae). Vol. 1. Text. Herbipoliana, 5 (1), 1–859.

    Berio, E. (1938 [1939]) Contributi alla conoscenza dei Lepidotteri Eteroceri dell’Eritrea. I. Lista delle specie con descrizioni delle nuove entità raccolte negli anni 1934 al 1937 dal Sig. Francesco Vaccaro. Memoire della Società Entomologica Italiana, 17, 47–62.

    Berio, E. (1940) Diagnosi di eteroceri africani. Memoire della Società Entomologica Italiana, 19, 125–128.

    Berio, E. (1954a) Note sulla sistematica dei generi Achaea Hb. e affini (Lep. Noctuidae) Bollettino della Societá Entomologica Italiana, 84, 22–24.

    Berio, E. (1954b) Osservacioni su Polydesma umbricola Bdv. e sul genere Polydesma Bdv. (Lep. Noctuidae) Doriana, 1 (50), 1–8.

    Berio, E. (1954 [1955]) Appunti sul genere Pericyma H.-S. e affini. Memoire della Società Entomologica Italiana, 33, 147–154.

    Berio, E. (1959) Studi sulla sistematica delle cosidette Catocalinae e Othreinae. Annali del Museo Civico di Storia Naturale Giacomo Doria, 71, 276–327.

    Berio, E. (1965) Le Catocaline africane a tibie spinose del Museo di Tervuren. Annali del Museo Civico di Storia Naturale Giacomo Doria, 75, 181–332.

    Berio, E. (1968) Revisione di alcune specie di Pandesma Guen. e Thria Wlk. con descrizione di nuove specie. Annali del Museo Civico di Storia Naturale Giacomo Doria, 77, 208–220.

    Berio, E. (1971) Revisione del gen. Polydesma Boisd. e Trichopolydesma Berio (Lepidoptera - Noctuidae). Annali del Museo Civico di Storia Naturale Giacomo Doria, 78, 264–300.

    Berio, E. (1988 [1989]) Novità in tema di Quadrifine paleartiche (Lepidoptera, Noctuidae). Bollettino della Società Entomologica Italiana, 120, 216–222.

    Berio, E. (1990) Descrizione di un nuovo genere di Ophiderini del Madagascar e di un nuovo genere di Cuculliinae del Perù. Bollettino della Società Entomologica Italiana, 122, 37–40.

    Berio, E. (1988 [1991]) La sistematica dell’antico genere neotropicale Letis Hbn. (e Blosyris Hbn.) (Lepidoptera, Noctuidae, Catocalinae, Ophiderini) con descrizione di 2 nuove specie. Frustula Entomologica (n. s.), 11, 31–40.

    Berio, E. (1991) Lepidoptera, Noctuidae. II. Sezione Quadrifide. Fauna D’Italia, Vol. 27. Edizioni Calderini, Bologna, xxv + 708 pp. + 16 plates.

    Berio, E. (1992) Ricerca di possibli modelli sistematici per le Catocalinae e Ophiderinae del globo. Memoire della Società Entomologica Italiana, 70, 287–303.

    Berio, E. & Fletcher, D.S. (1958) Monografia dell'antico genere Sypna Guen. (Lepidoptera – Noctuidae). Annali del Museo Civico di Storia Naturale Giacomo Doria, 70, 323–402.

    Berlinger, M.J., Yathom, S. & Halperin, J. (2001) Ophiusa tirhaca Cramer (Noctuidae: Lepidoptera) infesting pistachio trees in Israel. Zoology in the Middle East, 22, 83–86.
    http://dx.doi.org/10.1080/09397140.2001.10637851

    Börner, C. (1939) Die Grundlagen meines Lepidopterensystems. in Verhandlungen VII. Internationaler Kongress für Entomologie, 2, 1372–1424.

    Butler, A.G. (1892) Revision of the genus Melipotis Hübn., with descriptions of two new species. Annals and Magazine of Natural History. 6 (10), 315–327.
    http://dx.doi.org/10.1080/00222939208677417

    Cotes, E.C. & Swinhoe, C. (1888) Pt. III - Noctues, Pseudo-Deltoides, and Deltoides, In: Cotes, E.C. & Swinhoe, C. A catalogue of the moths of India. Trustees of the Indian Museum, Calcutta, pp. 256–461.

    Crumb, S.E. (1956) The larvae of the Phalaenidae. Technical Bulletin, United States Department of Agriculture, 1135, 356pp.

    Duponchel, P.A.J. (1844–1846) Catalogue méthodique des Lépidoptères d'Europe. Paris. 523 pp.

    Eggers, F. (1919) Das thoracale bitympanale Organ einer Gruppe der Lepidoptera Heterocera. Zoologische Jahrbücher, Abt. Anatomie und Ontogenie der Tiere, 41, 273–376.

    Fang, Q.Q., Mitchell, A., Regier, J.C., Mitter, C., Friedlander, T.P. & Poole, R.W. (2000) Phylogenetic utility of the nuclear gene Dopa Decarboxylase in noctuoid moths (Insecta: Lepidoptera: Noctuoidea). Molecular Phylogenetics and Evolution, 15, 473–486.
    http://dx.doi.org/10.1006/mpev.1999.0758

    Feige, R. (1977) Kurze Darstellung der neotropischen Gattung Letis Hbn. und ihrer Arten. Deutsche Entomologische Zeitschrift, 24, 123–139.
    http://dx.doi.org/10.1002/mmnd.19770240105

    Feige, K.R. (1988 [1991]) Drawings of the male genitalia of Letis species. Frustula Entomologica (n. s.), 11, 19–29.

    Fibiger, M. & Lafontaine, J.D. (2005) A review of the higher classification of the Noctuoidea (Lepidoptera) with special reference to the Holarctic fauna. Esperiana, 11, 7–92.

    Fletcher, D.S. & Nye, I.W.B. (1982) The generic names of moths of the World. Vol. 4. British Museum, Natural History, London, pp. 170.
    http://dx.doi.org/10.5962/bhl.title.119597

    Forbes, W.T.M. (1914) A table of the genera of Noctuidae of Northeastern North America. Journal of the New York Entomological Society, 22, 1–33.

    Forbes, W.T.M. (1954) The Lepidoptera of New York and Neighboring States. Part III. Noctuidae. Cornell University Agricultural Experiment Station Memoir, 329, 1–433.

    Franclemont, J.G. & Todd, E.L. (1983) Noctuidae. In: Hodges, R.W., Dominick, T., Davis, D.R., Ferguson, D.C., Franclemont, J.G., Munroe, E.G. & Powell, J.A. (Eds.), Check list of the Lepidoptera of North America North of Mexico. Classey, E.W. Ltd. and the Wedge Entomological Research Foundation, London, pp. 120–159.

    Fullard, J.H. & Napoleone, N. (2001) Diel flight periodicity and the evolution of auditory defenses in the Macrolepidoptera. Animal Behaviour, 62, 349–368.
    http://dx.doi.org/10.1006/anbe.2001.1753

    Gaede, M. (1939) Subfamily: Noctuinae. In: Seitz, A. (Ed.) (1934–1940) The Macrolepidoptera of the World, II. Division: The Macrolepidoptera of the African Fauna, XV. Volume: Noctuid Heterocera. iii + 286 pp. + 41 plates.

    Gardner, J.C.M. (1946a) On larvae of the Noctuidae (Lepidoptera) - I. Transactions of the Royal Entomological Society of London, 96, 61–72.
    http://dx.doi.org/10.1111/j.1365-2311.1946.tb00443.x

    Gardner, J.C.M. (1946b) On Larvae of the Noctuidae (Lepidoptera) - II. Transactions of the Royal Entomological Society of London, 97, 237–252.
    http://dx.doi.org/10.1111/j.1365-2311.1946.tb00265.x

    Gardner, J.C.M. (1947) On the larvae of the Noctuidae - III. Transactions of the Royal Entomological Society of London, 98, 59–89.
    http://dx.doi.org/10.1111/j.1365-2311.1947.tb01052.x

    Gardner, J.C.M. (1948a) Notes on pupae of the Noctuidae. Proceedings of the Royal Entomological Society of London, (B), 17, 84–92.
    http://dx.doi.org/10.1111/j.1365-3113.1948.tb00891.x

    Gardner, J.C.M. (1948b) On the larvae of the Noctuidae (Lepidoptera) - IV. Transactions of the Royal Entomological Society of London, 99, 291–318.
    http://dx.doi.org/10.1111/j.1365-2311.1948.tb01239.x

    Goater, B., Ronkay, L. & Fibiger, M. (2003) Noctuidae Europaeae Vol. 10, Catocalinae, Plusiinae. Entomological Press, Søro, 452 pp.

    Grote, A.R. (1874) List of the Noctuidae of North America. Bulletin of the Buffalo Society for Natural Science, 2, 1–77.

    Grote, A.R. (1889) The Noctuidae of North America and Europe compared, part I. The Canadian Entomologist, 21, 121–126.
    http://dx.doi.org/10.4039/Ent21121-7

    Grote, A.R. (1890a) The Noctuidae of North America and Europe compared, part VII. The Canadian Entomologist, 22, 105–109.
    http://dx.doi.org/10.4039/Ent22105-6

    Grote, A.R. (1890b) The Noctuidae of North America and Europe compared, part VIII. The Canadian Entomologist, 22, 145–149.
    http://dx.doi.org/10.4039/Ent22145-8

    Grote, A.R. (1895) The hypenoid moths and allied groups. Proceedings of the American Philosophical Society, 34, 416–436.

    Gueneé, A. (1841) Essai sur la classification des Noctuélides (suite et fin.). Annales de la Société Entomologique de France, 10, 217–250.

    Gueneé, A. (1852a) Noctuélites. In: Boisduval , J.A. and Guenée, A. (Eds.), Histoire Naturelle des Insectes - Species Général des Lépidoptères. Vol. 2. Roret, Paris, 444 pp.

    Gueneé, A. (1852b) Noctuélites. In: Boisduval , J.A. and Guenée, A. (Eds.), Histoire Naturelle des Insectes - Species Général des Lépidoptères. Vol. 3. Roret, Paris, 442 pp.

    Hacker, H.H. (2001) Fauna of the Nolidae and Noctuidae of the Levante with descriptions and taxonomic notes (Lepidoptera, Noctuoidea). Appendix: Revision of genus Clytie Hübner [1823]. Esperiana, 8, 7–398.

    Hampson, G.F. (1894) The fauna of British India, including Ceylon and Burma, Moths II. Taylor and Francis, London, 609 pp.

    Hampson, G.F. (1902) The moths of South Africa (Part II). Annals of the South African Museum of Natural History, 2, 255–446.

    Hampson, G.F. (1903) Catalogue of the Lepidoptera Phalaenae of the British Museum. Vol. 4, Trustees of the British Museum, London, 689 pp.

    Hampson, G.F. (1913a) Catalogue of the Lepidoptera Phalaenae of the British Museum. Vol. 12, Trustees of the British Museum, London, 626 pp.

    Hampson, G.F. (1913b) Catalogue of the Lepidoptera Phalaenae of the British Museum. Vol. 13, Trustees of the British Museum, London, 609 pp.

    Hemming, F. (1937) Hübner: A bibliographical and systematic account of the entomological works of Jakob Hübner (Vol. 1). Royal Entomological Society of London, London, 605 pp.

    Herrich-Schäffer, G.A.W. (1845) Systematische Bearbeitung der Schmetterlinge von Europa, zugleich lis Text, Revision und Supplement zu Jakob Hübner's Sammlung europäischer Schmetterlinge. Vol. 2. Die Schwärmer, Spinner und Eulen (Hepialides·Cossides·Zygaenides·Sesiides-Sphingides·Bombycides·Noctuides·Nycteolides) Vol. 2. J. Manz, Regensburg.

    Holloway, J.D. (2005) The Moths of Borneo (part 15 and 16): Family Noctuidae, Subfamily Catocalinae. Malayan Nature Journal, 58, 1–529.

    Holloway, J.D. (2011) The moths of Borneo: Families Phaudidae, Himantopteridae, and Zygaenidae; revised and annotated checklist. Malayan Nature Journal, 63, 1–548.

    Holloway, J.D. & Miller, S.E. (2003) The composition, generic placement, and host plant relationships of the joviana-group in the Parallelia genus complex (Lepidoptera: Noctuidae, Catocalinae). Invertebrate Systematics, 17, 111–128.
    http://dx.doi.org/10.1071/is02030

    Hübner, J. (1816 [1816–1826]) Verzeichniss bekannter Schmettlinge [sic.]. In: Augsburg, p. 431 pp. + 472 pp.

    John, O. (1909) A generic subdivision of the genus Palpangula Stgr. (Lepidoptera, Noctuidae). Revue Russe d'Entomologie, 9, 115–125.

    John, O. (1910) Eine Revision der Gattung Leucanitis Gn. Horae Societatis Entomologicae Rossicae, 39, 585–634.

    Kirby, W.F. (1897) Moths, Part III. In: A hand-book to the order Lepidoptera. Edward Lloyd Limited, London, pp. 1–199.
    http://dx.doi.org/10.5962/bhl.title.23911

    Kiriakoff, S.G. (1951) Recherches sur les organes tympaniques des Lépidoptères en rapport avec la classification. Entomologische Berichten, 13, 381–382.

    Kiriakoff, S.G. (1963) The tympanic structures of the Lepidoptera and the taxonomy of the order. Journal of the Lepidopterist's Society, 17, 1–6.

    Kitching, I.J. (1984) An historical review of the higher classification of the Noctuidae (Lepidoptera). Bulletin of the British Museum (Natural History), 49, 153–234.

    Kitching, I.J. & Rawlins, J. E. (1998) Evolution, Systematics and Biogeography. In: Kristensen (Ed.), Lepidoptera, Moths and Butterflies. Walter de Gruyter, Berlin, pp. 355–401.

    Klyuchko, Z.F. (1978) Fauna of Ukraine, Vol. 16: Cutworms; Quadrifine Complex: Nycteolinae, Hypeninae, Catocalinae, Othreinae, Euteliinae, Pantheinae, Plusiinae, Jaspidiinae. Naukova Dumka, Kiev, 415 pp.

    Фауна Укпаїни. (1978) Том 16. Совки Квадрифіноїдного Комплексу. Бипуск 6. З. Ф. Ключко. Ніктеоліни, Вусатки, Стрічкарки, Отреїни, Евтеліїни, Пантеїни, Металовидки, Яспідеїни. Київ «Наукова Думка», 415 с. Avaliable from: http://www.izan.kiev.ua/fau-ukr/vol16iss6.htm (Accessed 10 Nov. 2016)

    Kononenko, V.S. (2010) Noctuidae Sibiricae. Volume 2. Micronoctuidae, Noctuidae: Rivulinae - Agaristinae. Entomological Press, Sorø, 475 pp.

    Kononenko, V.S. & Pinratana, A. (2013) Moths of Thailand Vol. 3, Part 2. Noctuoidea. An illustrated Catalogue of Erebidae, Nolidae, Euteliidae, and Noctuidae (Insecta: Lepidoptera) in Thailand. Brothers of St. Gabriel in Thailand. Bangkok, 625 pp.

    Kozhantschikov, I.W. (1933) Ein neues Merkmal zur Begründung der Familien Trifidae und Quadrifidae. Deutsche Entomologische Zeitschrift “Iris”, Dresden, 47, 32–35.

    Kühne, L. (2005) Revision und Phylogenie der Gattungsgruppe Crypsotidia Rothschild, 1901, Tachosa Walker, 1869, Hypotacha Hampson, 1913, Audea Walker, [1858] 1857, und Ulotrichopus Wallengren, 1860 (Lepidoptera, Noctuidae, Catocalinae). Esperiana Memoir, 2, 7–220.

    Kühne, L. & Speidel, W. (2004) The system of the Catocalinae - a historical survey (Lepidoptera, Noctuidae). Beiträge zur Entomologie, 54, 147–184.

    Lafontaine, J.D. & Fibiger, M. (2006) Revised higher classification of the Noctuoidea (Lepidoptera). Canadian Entomologist, 138, 610–635.
    http://dx.doi.org/10.4039/n06-012

    Lafontaine, J.D. & Schmidt, B.C. (2010) Annotated checklist of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico. Zookeys, 40, 1–239.

    Lafontaine, J.D. & Schmidt, B.C. (2013) Additions and corrections to the check list of the Noctuoidea (Insecta, Lepidoptera) of North America north of Mexico. Zookeys, 264, 227–235.
    http://dx.doi.org/10.3897/zookeys.264.4443

    Lödl, M. (2000) The modification of the “posterior notal wing process” of the forewing in the family Noctuidae and its importance for taxonomy (Insecta, Lepidoptera). Quadrifina, 3, 303–323.

    Matov, A.Y. (2003) On the systematics of noctuid moths tribe Euclidiini sensu lato (Lepidoptera, Noctuidae, Catocalinae) of the Palaearctic. Entomologicheskoe Obozrenie, 82, 886–906.

    McDunnough, J.H. (1938) Check-list of the Lepidoptera of Canada and the United States of America. Part 1. Macrolepidoptera. Memoirs of the Southern California Academy of Sciences, 1, 1–272.

    Merzheevskaya, O.I. (1988) Larvae of owlet moths (Noctuidae): biology, morphology, and classification: Amerind Publ. Co., New Delhi, India, xx + 419 pp.

    Miller, J.S. (1991) Cladistics and classification of the Notodontidae (Lepidoptera: Noctuoidea) based on larval and adult morphology. Bulletin of the American Museum of Natural History, 204, 230 pp.

    Minet, J., Barbut, J. & Lalanne-Cassou, B. (2011) Les Noctuelles: classification et clef de détermination des familles (Lepidoptera: Noctuoidea). Alexanor, 25, 131–151.

    Mitchell, A., Cho, S., Regier, J.C., Mitter, C., Poole, R.W. & Matthews, M. (1997) Phylogenetic utility of Elongation Factor-1a in Noctuoidea (Insecta: Lepidoptera): The limits of synonymous substitution. Molecular Biology and Evolution, 14, 381–390.
    http://dx.doi.org/10.1093/oxfordjournals.molbev.a025774

    Mitchell, A., Mitter, C. & Regier, J.C. (2000) More taxa or more characters revisited: combining data from nuclear protein-encoding genes for phylogenetic analyses of Noctuoidea (Insecta: Lepidoptera). Systematic Biology, 49, 202–224.
    http://dx.doi.org/10.1093/sysbio/49.2.202

    Mitchell, A., Mitter, C. & Regier, J.C. (2006) Systematics and evolution of the cutworm moth (Lepidoptera: Noctuidae): evidence from two protein coding nuclear genes. Systematic Entomology, 31, 21–46.
    http://dx.doi.org/10.1111/j.1365-3113.2005.00306.x

    Mitter, C. (1988) Taxonomic potential of some internal reproductive structures in Catocala (Lepidoptera: Noctuidae) and related genera. Annals of the Entomological Society of America, 81, 10–18.
    http://dx.doi.org/10.1093/aesa/81.1.10

    Mitter, C. & Silverfine, E. (1988) On the systematic position of Catocala Schrank (Lepidoptera: Noctuidae). Systematic Entomology, 13, 67–84.
    http://dx.doi.org/10.1111/j.1365-3113.1988.tb00230.x

    Moore, F. (1884–1887) The Lepidoptera of Ceylon (Vol. 3). Reeve, L. & Co., London.

    Mosher, E. (1916) A classification of the Lepidoptera based on characters of the pupa. Bulletin of the Illinois State Laboratory of Natural History, 12, 17–159.

    Muniappan, R. (1974) Biology of the Poinciana Looper, Pericyma cruegeri (Butler) on Guam. Micronesica, 10, 273–278.

    Naik, M.I., Ajith Kumar, M.A., Manjunatha, M. & Shivanna, B.K. (2010) Survey for the pests of castor and natural enemies of castor semilooper. Environment and Ecology (Kalyani), 28, 558–563.

    Ogunwolu, E.O. & Habeck, D.H. (1975) Comparative life histories of three Mocis spp. in Florida, USA (Lepidoptera: Noctuidae). Florida Entomologist, 58, 97–103.
    http://dx.doi.org/10.2307/3493389

    Olivares, T.S. & Angulo, A.O. (1996) El organo timpanico en la clasificacion de Lepidoptera: Noctuidae. Boletín de Entomología Venezolana. 11, 155–183.

    Packard, A.S. (1869) The characters of the lepidopterous family Noctuidae. Proceedings of the Portland Society of Natural History, 1, 153–156.

    Pagenstecher, A. (1894) Beiträge zur Lepidopteren-Fauna des malayischen Archipels (IX), 1. Über Javanische Schmetterlinge. Jahrbücher des Nassauischen Vereines für Naturkunde, 47, 27–51.

    Pavesi, F., de Vos, R. & Zilli, A. (2010) Revision of the genus Ommatophora Guenée, 1852 with description of four new species (Lepidoptera: Noctuidae). Quadrifina, 9, 483–510.

    Prout, A.E. (1921) Notes on some Noctuidae in the Joicey collection, with descriptions of new species. The Annals and Magazine of Natural History, 8, 1–33.
    http://dx.doi.org/10.1080/00222932108632557

    Richards, A.G. (1932 [1933]) Comparative skeletal morphology of the noctuid tympanum. Entomologica Americana, 13, 1–43, pl. 41–20.

    Richards, A.G. (1935a) Forsebia - A new genus of American Erebinae (Lepidoptera, Noctuidae). Canadian Entomologist, 67, 264–267.
    http://dx.doi.org/10.4039/Ent67264-12

    Richards, A.G. (1935b) Notes on the structure and position of Drasteriodes Hampson (Lepid.: Noctuidae). Entomological News, 46, 129–141.

    Richards, A.G. (1936) A generic synopsis of the Phoberia-Melipotis-Drasteria-Boryza series of Erebinae (Lepidoptera: Noctuidae). Revista de Entomologia Rio de Janeiro, 6, 338–374.

    Richards, A.G. (1939) A revision of the North American species of the Phoberia-Melipotis-Drasteria group of moths (Lepidoptera: Phalaenidae). Entomologica Americana, 19, 1–100.

    Roepke, W. (1941) On some new or little known Indo-Malayan noctuids (Lepidoptera Heterocera, family Agrotidae) Zoologische Mededelingen, 23, 13–20.

    Rota, J., Zacharczenko, B.V., Wahlberg, N., Zahiri, R., Schmidt, B.C. & Wagner, D.L. (2015) Phylogenetic relationships of Acronictinae with discussion of the abdominal courtship brush in the Noctuidae (Lepidoptera). Systematic Entomology, 41, 416–429.
    http://dx.doi.org/10.1111/syen.12162

    Ryabov, M.A. (1988) Review of the morphology of noctuid moths (Lepidoptera: Noctuidae). In Kryzhanovskii, O.L. (Ed.) Lepidopterous Fauna of the USSR and Adjacent Countries, Oxonian Press, New Delhi. pp. 348–387.

    Sargent, T.D. (1976) Legion of Night: The Underwing Moths. University of Massachusetts Press, Amherst. xii + 222 pp. + 8 plates.

    Seitz, A. (1914) Preface. In: Seitz, A. (Ed.) (1909–1914). The Macrolepidoptera of the World. The Macrolepidoptera of the Palearctic Region, III. Volume: The Noctuid Moths. Alfred Kernen, Stuttgart. iv + 511 pp. + 50 plates.

    Smith, J.B. (1882a) Synopsis of the North American genera of Noctuidae. Bulletin of the Brooklyn Entomological Society, 5, 33–36.

    Smith, J.B. (1882b) Synopsis of the North American genera of Noctuidae. Bulletin of the Brooklyn Entomological Society, 5, 27–30.

    Smith, J.B. (1908) A revision of some species of Noctuidae heretofore referred to the genus Homoptera Boisduval. Proceedings of the United States National Museum, 35, 209–275 + plates 31–36.
    http://dx.doi.org/10.5479/si.00963801.35-1645.209

    Speidel, W., Fänger, H. & Naumann, C.M. (1996a) The phylogeny of the Noctuidae. Systematic Entomology, 21, 219–251.
    http://dx.doi.org/10.1046/j.1365-3113.1996.d01-14.x

    Speidel, W., Fänger, H. & Naumann, C.M. (1996b) The surface microstructure of the noctuid proboscis (Lepidoptera: Noctuidae). Zoologischer Anzeiger, 234, 307–315.

    Speidel, W., Fänger, H. & Naumann, C.M. (1997) On the systematic position of Cocytia Boisduval, 1828 (Lepidoptera: Noctuidae). Deutsche Entomologische Zeitschrift, 44, 27–31.
    http://dx.doi.org/10.1002/mmnd.19970440104

    Speidel, W. & Naumann, C.M. (1995a) Further morphological characters for a phylogenetic classification of the Noctuidae (Lepidoptera). Beiträge zur Entomolgie, 45, 119–135.

    Speidel, W. & Naumann, C.M. (1995b) Phylogenetic aspects in the higher classification of the subfamily Catocalinae (Lepidoptera, Noctuidae). Beiträge zur Entomologie, 45, 109–118.

    Speidel, W. & Naumann, C.M. (2004) A survey of the family group names in noctuoid moths (Insecta: Lepidoptera). Systematics and Biodiversity, 2, 191–221.
    http://dx.doi.org/10.1017/S1477200004001409

    Stekolnikov, A.A. & Speidel, W. (2009) Taxonomische Stellung der Gattungen Panthea, Trichosea, und Diloba (Lepidoptera, Noctuoidea, „Pantheidae” und Noctuidae). Entomofauna, 30, 61–104.

    Sugi, S. (1968) Melapia new genus, with description of a new species from the Ryukyus (Loochoos) (Lepidoptera, Noctuidae, Catocalinae). Kontyû, 36, 164–168.

    Tikhomirov, A.M. (1979) The system and phylogeny of the palaearctic Noctuidae (Lepidoptera) as indicated by the functional morphology of the male genitalia. Entomological Review, 58, 119–133.

    Tutt, J.W. (1896) British Moths. Routledge and Sons, London, 368 pp.

    van Nieukerken, E.J, Kaila, L., Kitching, I.J., Kristensen, N.P., Lees, D.J., Minet, J., Mitter, J., Mutanen, M., Regier, J.C., Simonsen, T.J., Wahlberg, N., Yen, S-H., Zahiri, R., Adamski, D., Baixeras, J., Bartsch, D., Bengtsson, B.Å., Brown, J.W., Bucheli, R.S., Davis, D.R., De Prins, J., De Prins, W., Epstein, M.E., Gentili-Poole, P., Gielis, C., Hättenschwiler, P., Hausmann, A., Holloway, J.P., Kallies, A., Karsholt, O., Kawahara, A., Koster, S.J., CKozlov, M.V., Lafontaine, J.D., Lamas, G., Landry, J-F., Lee, S., Nuss, M., Park, K-T., Penz, C., Rota, J., Schmidt, B.C., Schintlmeister, A., Sohn, J.C., Solis, M.A., Tarmann, G., Warren, A.D., Weller, S., Yakovlev, Y., Zolotuhin, V.V. & Zwick, A. (2011) Order Lepidoptera Linnaeus, 1758. In: Zhang Z-Q. (Ed.) Animal biodiversity: An outline of higher-level classification and survey of taxonomic richness. Zootaxa, 3148, 212–221.

    Vázquez, Y.M., Martínez, A.M., Valdez, J.M., Figueroa, J.I., Chavarrieta, J.M., Rebollar, A., Sánchez, J.A. & Viñuela y S. Pineda, E. (2014) Life History, Diagnosis, and Parasitoids of Zale phaeograpta (Lepidoptera: Erebidae). Annals of the Entomological Society of America, 107, 170–177.
    http://dx.doi.org/10.1603/AN13093

    Wagner, D.L., Schweitzer, D.F., Sullivan, J.B. & Reardon, R.C. (2011) Owlet Caterpillars of Eastern North America (Lepidoptera: Noctuidae): Princeton University Press, Princeton, NJ, 576 pp.

    Walker, F. (1857) List of the specimens of lepidopterous insects in the collection of the British Museum. Part XII.--Noctuidae. Trustees of the British Museum, London, pp. 766–982.

    Walker, F. (1857) List of the specimens of lepidopterous insects in the collection of the British Museum. Part XIII.--Noctuidae. Trustees of the British Museum, London, pp. 983–1236.

    Walker, F. (1858) List of the specimens of lepidopterous insects in the collection of the British Museum. Part XIV.--Noctuidae. Trustees of the British Museum, London, pp. 1237–1519.

    Walker, F. (1858) List of the specimens of lepidopterous insects in the collection of the British Museum. Part XIII.--Noctuidae, Trustees of the British Museum, Natural History, London, pp. 1520–1888.

    Weller, S.J., Friedlander, T.P., Martin, J.A. & Pashley, D.P. (1992) Phlyogenetic studies of ribosomal RNA variation in higher moths and butterflies (Lepidoptera: Ditrysia). Molecular Phylogenetics and Evolution, 1, 312–337.
    http://dx.doi.org/10.1016/1055-7903(92)90007-4

    Weller, S.J., Pashley, D.P., Martin, J.A. & Constable, J.L. (1994) Phylogeny of noctuoid moths and the utility of combining independent nuclear and mitochondrial genes. Systematic Biology, 43, 194–211.
    http://dx.doi.org/10.2307/2413461

    Wiltshire, E.P. (1970) Middle East Lepidoptera XVIII, A review of the genus Pericyma Herrich-Schäffer and neighbouring genera, and especially their relationships as shown by their genitalia, with a description of a new species from Abyssinia,. Veröffentlichungen der Zoologischen Staatssammlung München, 14, 91–119.

    Wiltshire, E.P. (1976) Middle East Lepidoptera, XXXII, Diagnosis of some eremic tribes of Noctuidae - Quadrifinae, with a discussion of their biogeographical significance. Journal of the Bombay Natural History Society, 73, 157–165.

    Wiltshire, E.P. (1990) An illustrated, annotated catalogue of the Macro-Heterocera of Saudi Arabia. Fauna of Saudi Arabia, 11, 91–250.

    Yagi, N. & Koyama, N. (1963) The Compound Eye of the Lepidoptera: An Approach from Organic Evolution. Shinkyo Press Co., Ltd., Tokyo, 323 pp.

    Yela, J.L. (1997) Noctuidos del área Iberobalear: Adiciones y correciones a la lista sistemática, con consideraciones micro y macroevolutivas y una propuesta filogenética global (Insecta: Lepidoptera: Noctuidae). Zapateri, 7, 91–190.

    Yela, J.L. & Kitching, I.J. (1999) Noctuid phylogeny revisited (Insecta: Lepidoptera: Noctuidae). Boletín de la Sociedad Entomológica Aragonesa, 26, 485–520.

    Zahiri, R., Holloway, J.D., Kitching, I.J., Lafontaine, J.D., Mutanen, M. & Wahlberg, N. (2012) Molecular phylogenetics of the Erebidae (Lepidoptera, Noctuoidea). Systematic Entomology, 37, 102–124.
    http://dx.doi.org/10.1111/j.1365-3113.2011.00607.x

    Zahiri, R., Kitching, I.J., Lafontaine, J.D., Mutanen, M., Kaila, L., Holloway, J.D. & Wahlberg, N. (2010) A new molecular phylogeny offers hope for a stable family level classification of the Noctuoidea (Lepidoptera). Zoologica Scripta, 40, 158–173.
    http://dx.doi.org/10.1111/j.1463-6409.2010.00459.x

    Zahiri, R., Lafontaine, J.D., Schmidt, B.C., Holloway, J.D., Kitching, I.J., Mutanen, M. & Wahlberg, N. (2013) Relationships among the basal lineages of Noctuidae (Lepidoptera, Noctuoidea) based on eight gene regions. Zoologica Scripta, 42, 488–507.
    http://dx.doi.org/10.1111/zsc.12022

    Zilli, A. (2003) A new species of the late genus Letis Hübner, [1821] from Ecuador: an exercise of symmetry (Lepidoptera: Noctuidae). Quadrifina, 6, 141–152.