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Type: Monograph
Published: 2016-03-14
Page range: 1–81
Abstract views: 58
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Annotated checklist of Georgian oribatid mites 

Institute of Entomology. Agricultural University of Georgia. 240, D. Aghmashenebely Alley 0131. Tbilisi, Georgia.
Institute of Ecology of Ilia State University. Cholokashvili ave 3/5, 0165, Tbilisi, Georgia. Invertebrate Research Centre (IRC), Agladze Str. 26, 0119 Tbilisi, Georgia
Hemiptera Oribatid mites catalogue Caucasus biodiversity

Abstract

A new updated checklist of Georgian oribatid mites is based on the critical review of existing literature data and new findings. The list includes 534 oribatid species of which 21 species are new for the country recorded from more than 390 locations. For each species information of the global and regional distribution is presented with notes on ecological characteristics. As far as necessary we provide remarks on taxonomic issues to overcome the ambiguities and inconsistencies existing in literature. 

 

References

  1. Aoki, J. (1995) Oribatid mites of high altitude forests of Taiwan. Special Bulletin of Japanese Society of Coleopterologists, 4, 123–130.

    Arabuli, T., Murvanidze, M. & Kvavadze, Er. (2004) Distribution of oribatid mites (Acari, Oribatida) by phytocenoses (Tsiv-Gombori Range, East Georgia). Proceedings of the institute of zoology, XXII, 80–88.

    Arabuli, G., Mosulishvili, M., Murvanidze, M., Arabuli, T., Bagaturia, N. & Kvavadze, Er. (2007) The Colchic lowland alder woodland with boxwood understorey (Alneta barbatae buxosa) and their soil invertebrate animals. Proceedings of Georgian Academy of Sciences, Seria Biologia, 5 (2), 35–42

    Arabuli, G., Kvavadze, E., Kikodze, D., Connor, S.E., Kvavadze, Er., Bagaturia, N., Murvanidze, M. & Arabuli, T. (2008) The Krummholz beech woods of Mt. Tavkvetili (Javakheti plateau, Southern Georgia) – a relict ecosystem. Proceedings of the Institute of Zoology, XXIII, 194–213.

    Barjadze, Sh. & Murvanidze, M. (2016) New records of the springtails (Collembola: Entomobryomorpha) and oribatid mites (Acari: Oribatida) in Georgia. Turkish Journal of Zoology, 40, 117–119.
    http://dx.doi.org/10.3906/zoo-1502-8

    Behan-Pelletier, V.M. & Eamer, B. (2007) Aquatic oribatida: adaptations, constrains, distribution and ecology. In: Morales-Maracala, J.B., Behan-Pelletier,V., Ueckermann, E., Perez, T.M., Estrada-Venegas, E.G & Badii, M. (Eds.), Acarology. XI. Proceedings of the 11th International Congress of Acarology. Instituto de Biología and Facultad de Ciencias, Universidad Nacional Autónoma de México; Sociedad Latinoamericana de Acarología. México, Merida, pp. 71–82.

    Behan-Pelletier, V.M. & Eamer, B. (2008) Mycobatidae (Acari: Oribatida) of North America. Canadian Entomologist, 140, 73–110.
    http://dx.doi.org/10.4039/n07-027

    Bernini, F., Castagnoli, M. & Nannelli, R. (1995) Arachnida, Acari. In: Minelli, A., Ruffo, S. & La Posta, S. (Eds.), Checklist delle specie della fauna Italiana. Commission of the European Communities. Calderini, Bologna, 131 pp.

    Bulanova-Zakhvatkina, E.M. (1970) Oribatid mite fauna of the USSR and their distribution. In: Bulanova-Zakhvatkina, E.M., Ghilarov M.S., Krivolutsky, D.A., Petrova-Nikotina, A.D., Eitminavichute, I.S. & Aukshtikalnene A.M. (Eds), Oribatid mites and their role in the process of the soil formation. Akademia Nauk Litovskoi SSR, Vilnus, pp. 55–71. [in Russian]

    Colloff, M.J. & Cameron, S.L. (2013) A phylogenetic analysis and taxonomic revision of the oribatid mite family Malaconothridae (Acari: Oribatida), with new species of Tyrphonothrus and Malaconothrus from Australia. Zootaxa, 3681 (4), 301–346.

    http://dx.doi.org/10.11646/zootaxa.3681.4.1

    Darejanashvili, Sh. (1964) To the study of species composition of oribatid mites (Acari, Oribatei) in Tbilisi surroundings. Bulletin of the Academy of Sciences of the Georgian SSR, XXXIV (2), 457–464. [in Georgian]

    Darejanashvili, Sh. (1967) On the distribution of oribatid mites of Borjom-Kharagauli gorge. Bulletin of the Academy of Sciences of the Georgian SSR, XIV (3), 721–726. [in Russian]

    Darejanashvili, Sh. (1976) Oribatid mites (Oribatei) of Eastern Georgia. Summary of the PhD thesis, Metsniereba, Tbilisi, 24 pp. [in Russian]

    Darejanashvili, Sh. (1987) Oribatid mties of lowhill-steppe landscapes. In: Kurashvili, B. (Ed.), Problemi pochvennoi zoologii. Tezisi dokladov IX Vsesoyuznogo soveshchaniya, Metsniereba, Tbilisi, pp. 82–83. [in Russian]

    Darejanashvili, Sh. (2000) Oribatid mites of the coniferous and broadleaved forests of Trialeti Range. Proceedings of the Institute of Zoology, XX, 100–108. [in Georgian]

    Darejanashvili, Sh. & Gurgenidze, L. (2004) Ecological complexes of the oribatid mites (Acari, Oribatei) in the different forest types of the Eastern Georgia. Proceedings of the institute of Zoology, XXII, 102–110. [in Russian]

    Djaparidze, N. (1963) To the study the oribatid mite (Acari, Oribatei) fauna from Georgia. Bulletin of the Academy of Sciences of the Georgian SSR, XXXI (2), 413–419. [in Georgian]

    Djaparidze, N. (1966) Oribatid mites of Trialeti range – habitat distribution of the fauna. In: Reck, H.F. (Ed.), Invertebrate Fauna of the Trialetsky Mountains. Metsniereba, Tbilisi, pp. 40–63 [in Russian]

    Djaparidze, N. (1973) The description of two new species of oribatid mties (Acarina, Oribatei) from Georgia. Bulletin of the Academy of Sciences of Georgian SSR, 71 (2), 469–472. [in Russian]

    Djaparidze, N. (1974) Oribatid mites of the several natural zones of Georgia. In: Reck, H.F. (Ed.), Materiali k faune Gruzii. 4. Metsniereba, Tbilisi, pp. 11–40 [in Russian]

    Djaparidze, N. (1979) Oribatid mites of steppe biotopes of Martkopi surroundings. In: Reck, H.F. (Ed.), The invertebrates of brown and montane black soils of Georgia. Metsniereba, Tbilisi, pp. 157–166. [in Russian]

    Djaparidze, N. (1985) New findings of oribatid mites (Acariformes, Oribatei) in Georgia. In: Cholokava, A.O. (Ed.), Fauna i ekologia nekotorikh grupp nasekomikh i kleshchei Gruzii. Metsniereba. Tbilisi, Tbilisi, pp. 58–71. [in Russian]

    Djaparidze, N. (1989) Oribatid mites (Oribatei) in Georgian fauna. In: Eliashvili, T. (Ed.), Fauna i ekologia bezpozvonochnikh zhivotnikh Gruzii, Metsniereba, pp. 65–70. [in Russian]

    Djaparidze, N. (1990) New species of oribatid mites (Oribatei) of genus Flexa in Georgian fauna. Bulletin of the Academy of Sciences of the Georgian SSR, 137 (2), 405–408. [in Russian]

    Djaparidze, M. (1990a) Two new species of oribatid mites (Oribatei) of Oribatula Berlese, 1896 and Eremaeus C.L. Koch, 1836 genera in the fauna of Georgia. Bulletin of the Academy of Sciences of Georgian SSR, 139 (2), 417–420. [in Russian]

    Dubinina, E.V., Sosnina, E.F., Visotskaya, S.O., Markov, G.N. & Atanasov, L. Kh. (1966) Oribatid mties (Oribatida) from rodent nests on the Vitosh Mountain. Bulletin de L’Institut de Zoologie et Musee, XXII, 81–141. [in Russian]

    Engelbrecht, C.M. (1972) Ctenogalumna moresonensis sp.n. and Pilogalumna bloemfonteinensis sp.n., two new South African species of the subfamily Allogalumnidae Balogh, 1960 (Galumnidae: Oribatei). Acarologia, XIV (3), 497–510.

    Ermilov, S., Shtanchaeva, U. Ya., Subías, L.S. & Martens, J. (2014) Two new species of oribatid mites of Lasiobelba (Acari, Oribatida, Oppiidae) from Nepal, including a key to all species of the genus. ZooKeys, 424, 1–17.
    http://dx.doi.org/10.3897/zookeys.424.7990

    Ghilarov, M.S. & Krivolutsky, D.A. (1975) Identification keys of soil inhabiting mites, Sarcoptiformes. Nauka, Moscow, 491 pp. [in Russian]

    Grobler, L., Bayram, S. & Çobanoglu, S. (2004) Two new species and new records of oribatid mites from Turkey. International Journal of Acarology, 30 (4), 351–358.
    http://dx.doi.org/10.1080/01647950408684405

    Hsieh, T.C., Ma, K.H. & Chao, A. (2013) iNEXT online: interpolation and extrapolation (Version 1.0) [Software]. Available from http://chao.stat.nthu.edu.tw/blog/software-download/ (accessed 1 December 2015)

    Iordansky, S.N. (1991) Revision of oribatid mites of genus Oribatula (Acariformes, Cryptostigmata, Oribatulidae) of the fauna of USSR. Zoologichesky Zhurnal, 70 (8), 77–89. [in Russian]

    Karppinen, E., Krivolutsky, D.A., Tarba, Z.M., Shtanchaeva, U. Ya. & Gordeeva, E.W. (1987) List of oribatid mites (Acarina, Oribatei) of northern palaearctic region. IV. Caucasus and Crimea. Annales Entomologici Fennici, 53, 119–137.

    Krisper, G. (1984) Wiederbeschreibung und Verbreitungsanalyse der bodenbewohnenden Milbe Zetorchestes falzonii Coggi (Acari, Oribatei). Mitteilungen naturwissenschaftlichen Verlag Steiermark, 127, 147–152.

    Krivolutsky, D.A. (1975) Family Liacaridae. In: Ghilarov, M.S. & Krivolutsky, D.A. (Eds.), Identification keys of soil inhabiting mites, Sarcoptiformes. Nauka, Moscow, pp. 174–181. [in Russian]

    Krivolutsky, D.A. & Tarba, Z.M. (1971) On the fauna of oribatid mites of Abkhazia. Zoologichesky Zhurnal, 50, 1408–1411. [in Russian]

    Krivolutsky, D.A. & Tarba, Z.M. (1972) The fauna of oribatid mites of Abkhazia. In: Krivolutsky, D.A. (Ed.), Ekologia pochvennikh bezpozvonochnikh, Nauka, Moscow, pp. 203–207. [in Russian]

    Kulijev, K.A. (1961) To the study of oribatid mties of Azerbaijan with the description of two new species. Trudi Azerbaijanskogo Gosudarstvennogo universiteta, XVII, 47–58. [in Russian]

    Lange, A.B. (1972) New oribatid mites of the family Palaeacaridae. Vestnik MGU. Seria Biologia, 4, 103–106. [in Russian]

    Lange, A.B. (1972a) New species of Genus Zachvatkinella (Palaeacaridae, Acariformes) from Far East, Baykal and Caucasus. Zoologichesky Zhurnal, 51 (12), 1889–1892. [in Russian]

    Mahunka, S. (1979) Neue und interessante Milben aus dem Genfer Museum. XLI. Vierter Beitrag der Oribatiden-Fauna Griechenlands (Acari, Oribatida). Revue Suisse de Zoologie, 86 (2), 541–571.
    http://dx.doi.org/10.5962/bhl.part.82319

    Mahunka, S. (1982) Neue und interessante Milben aus dem Genfer Museum XXXIX. 1 Fifth contribution to the oribatid fauna of Greece. Revue Suisse de Zoologie, 89 (2), 497–515.
    http://dx.doi.org/10.5962/bhl.part.82456

    Mahunka, S. (1991) Notes, additions and redescriptions of the oribatid species of Berlese (Acari). Acta Zoologica Hungarica, 37 (1–2), 27–58.

    Mahunka, S. (1992) “Pelops” and “Oribates” species in the Berlese-collection (Acari). Acta Zoologica Hungarica, 38 (3–4), 213–260.

    Mahunka, S. & Mahunka-Papp, L. (2004) A catalogue of Hungarian oribatid mites. In: Cszuzdi, C. & Mahunka, S. (Eds.), Pedozoologica Hungarica. 2. Hungarian Natural History Museum and Systematic Zoology Research Group of the Hungarian Academy of Sciences, Budapest, 363 pp.

    Magurran, A.E. (2004) Measuring Biological Diversity. Blackwell Publishing, Cornwall, 247 pp.

    Maruashvili, M. (1964) Physical geography of Georgia. Metsniereba, Tbilisi, 343 pp. [in Georgian]

    Mumladze, L., Murvanidze, M. & Behan-Pelletier, V. (2013) Compositional patterns in Holarctic peat bog inhabiting oribatid mite (Acari: Oribatida) communities. Pedobiologia, 56, 41–48.
    http://dx.doi.org/10.1016/j.pedobi.2012.10.001

    Mumladze, L., Murvanidze, M., Maraun, M. & Salakaia, M. (2015) Oribatid mite communities along an elevational gradient in Sairme Gorge (Caucasus). Experimental and Applied Acarology, 66 (1), 41–51.
    http://dx.doi.org/10.1007/s10493-015-9893-4

    Murvanidze, M. (2000) The list of oribatid mites of Tbilisi city. Proceedings of the institute of zoology, XX, 109–118. [in Georgian]

    Murvanidze, M. (2002) Zoogeographical distribution of oribatid mites (Acari, Oribatei) of Tbilisi. Bullettin of Georgian Academy of Sciences, 165 (2), 371–373.

    Murvanidze, M. (2014) Oribatid mites of Georgian (Caucasus) caves including the description of a new species of Ghilarovus Krivolutsky, 1966. International Journal of Acarology, 40 (6), 463–474.
    http://dx.doi.org/10.1080/01647954.2014.950604

    Murvanidze, M. & Darejanashvili, Sh. (2000) Checklist of oribatid mites of Georgia. Proceedings of the institute of zoology, XX, 119–137.

    Murvanidze, M., Kvavadze, E. & Jgenti, L. (2004) Oribatid mites (Acari, Oribatei) of Ajara (Caucasus, Georgia) and their vertical-zonal distribution. Proceedings of the Institute of Zoology, XXII, 89–102.

    Murvanidze, M. & Kvavadze, E. (2006) Bioindication of semidesert, steppe and light forest oribatid mites (Acari, Oribatida) communities via isovalent species groups. Proceedings of Georgian Academy of Sciences. Biology, Seria B, 4 (4), 51–56.

    Murvanidze, M. & Kvavadze, Er. (2007) Oribatid mites (Acari, Oribatida) of the Lagodekhi Reserve. In: Lebedeva, N. (Ed.), Studies of the Southern Scientific Centre of the Russian Academy of Sciences. Issue III. Biodiversity and transformation of mountain ecosystems of Caucasus. SSC RAS Publishing, Rostov-on-Don, pp. 124–130. [in Russian]

    Murvanidze, M. & Kvavadze, Er. (2009) An inventory of oribatid mites, the main decomposers in bogs of Colchic Lowland (Caucasus, Georgia). In: Sabelis, M.W. & Bruin, J. (Eds.), Trends in Acarology, Springer, Dordrecht/Heidelberg/New York/London, pp. 175–178.

    Murvanidze, M. & Weigmann, G. (2003) Contribution to the oribatid mite fauna of Georgia. 1. New species of Poronota (Acari, Oribatida). Spixiana, 26 (2), 165–170.

    Murvanidze, M. & Weigmann, G. (2007) New Carabodidae (Acari: Oribatida) of Georgia. Tijdschrift voor Entomologie, 150, 193–200.
    http://dx.doi.org/10.1163/22119434-900000219

    Murvanidze, M., Arabuli, T., Bagathuria, N., Eliava, I., Kvavadze, Er. & Mumladze L. (2008) The nematodes and oribatid mites as indicators of urban environment. Proceedings of the Institute of Zoology, XXIII, 180–193.

    Murvanidze, M., Kvavadze, Er. & Jgenti, L. (2008a) Oribatid mites (Acari, Oribatida) of Castanea forests of Mtirala National Park and Kintrishi Reserve. Proceedings of Georgian Academy of Sciences. Biology, Seria B, 6 (1–2), 17–22.

    Murvanidze, M. & Behan-Pelletier, V.M. (2011) A new species of Striatoppia (Acari, Oribatida) from the Caucasus region, with remarks on the familial placement of the genus. International Journal of Acarology, 37 (1), 53–59.
    http://dx.doi.org/10.1080/01647954.2010.494165

    Murvanidze, M., Mumladze, L., Arabuli, T. & Kvavadze, Er. (2011) Landscape distribution of oribatid mites (Acari, Oribatida) in Kolkheti National Park (Georgia, Caucasus), Zoosymposia, 6, 221–233.

    Murvanidze, M. & Weigmann G. (2012) Two new species of oribatid mites (Acari, Oribatida) Haplozetes longisacculus and Scutovertex armazi from Georgia (Caucasus). Acarina, 20 (2), 167–172.

    Murvanidze, M., Mumladze, L., Arabuli, T. & Kvavadze, Er. (2013) Oribatid mite colonization of sand and manganese tailing sites. Acarologia, 53 (2), 127–139.
    http://dx.doi.org/10.1051/acarologia/20132089

    Murvanidze, M. & Mumladze, L. (2014) Oribatid mite (Acari, Oribatida) diversity in different forest stands of Borjom-Kharagauli National Park. Persian Journal of Acarology, 3 (4), 257–276.

    Murvanidze, M. & Arabuli, T. (2015) Oribatid mite diversity in Rhododendron ponticum L. canopy along elevational gradient in Mtirala National Park. Acarologia, 55 (2), 219–30.
    http://dx.doi.org/10.1051/acarologia/20152162

    Murvanidze, M., Mumladze, L., Arabuli, T., Barjadze, Sh. & Salakaia, M. (2016) Oribatida diversity in different microhabitats of Mtirala National Park. Journal of Japanese Association of Acarologists. [in press]

    Murvanidze, M. & Todria, N. (2015) Oribatida diversity on limestone and clay quarries. Proceedings of the Institute of Zoology, 24, 159–169.

    Niedbała, W. (1983) Deux neuveaux Phthiracaridae (Acarida, Oribatida) de la Georgie (URSS). Bullettin de la societe des amis des sciences et des lettres de Poznan, 23, 171–182.

    Niedbała, W. (1983a) Les nouveaux Phthiracaridae (Acari, Oribatida) du Caucase. Annales Zoologici, 37 (1), 1–62.

    Niedbała, W. (1986) Catalogue des Phthiracaroiea (Acari), clef pour la determination des especes et descriptions d’especes nouvelles. Annales Zoologici, 40 (4), 309–370.

    Niedbała, W. (1993) Revision of oribatid mites of Berlese’s collection. III. Redescription of species from Mesoplophoroidea and Euphthiracaroidea (Acari, Oribatida). Genus, 4 (1), 41–58.

    Niedbała, W. (2002) Ptyctimous mites (Acari, Oribatida) of the Nearctic region. In: Holeksa, K. (Ed.), Monographs of the Upper Silesian Museum, 4, pp. 1–261. [Bytom]

    Niedbała, W. (2004) Ptyctimous mites (Acari, Oribatida) of the Neotropical region. Annales Zoologici, 54 (1), 1–288.

    Niedbała , W. (2011) Ptyctimous mites (Acari: Oribatida) of the Palaearctic Region. Systematic part. In: Ivan, D. & Maķol, J. (Eds.), Fauna Mundi. 4. Natura optima dux Foundation, Warszawa, 472 pp.

    Niedbała, W. (2015) Current taxonomical and faunistic status of Caucasian ptyctimous mites (Acari, Oribatida). North-Western Journal of Zoology, 11 (1), 1–7.

    Norton, R.A. (1994) Evolutionary aspects of oribatid mite life histories and consequences for the origin of Astigmata. In: Houk, M. (Ed.), Ecological and evolutionary analyses of life history patterns. Chapman & Hall, New York, pp. 99–135.
    http://dx.doi.org/10.1007/978-1-4615-2389-5_5

    Norton, R.A. & Behan-Pelletier, V.M. (2009) Suborder Oribatida. In: Walter & Kranz (Eds.), A manual of Acarology. Third edition. Texas Technical University Press, Lubbock, pp. 430–564.

    Norton, R.A. & Sidorchuk, E.A. (2014) Collohmannia johnstoni n.sp. (Acari, Oribatida) from West Virginia (U.S.A.), including description of ontogeny, setal variation, notes on biology and systematics of Collohmanniidae. Acarologia, 54 (3), 271–334.
    http://dx.doi.org/10.1051/acarologia/20142134

    Ohkubo, N. (1995) Species list of Quadroppiidae (Acari: Oribatida) with descriptions of a new genus and two new species. Journal of the Acarological Society of Japan, 4 (2), 77–89.
    http://dx.doi.org/10.2300/acari.4.77

    Oudemans, A.C. (1914) Acarologisches aus Maulwurfsnestern. Archiv fuer Naturgeschichte, A (10), 1–70.

    Perez-Iñigo, C. (1993) Acari, Oribatei, Poronota. In: Ramos Sanches, M.A. (Ed.), Fauna Iberica. 3, Museo Nacional de Ciencias Naturales, CSIC, Madrid, 320 pp.

    Reck, H. (1976) The Catalogue of the Acarofauna of Georgian SSR. Metsniereba, Tbilisi, 128 pp. [in Russian]

    Schatz, H. (2004) Hornmilben (Acari, Oribatida) in Auenwäldern an der Etsch und Tafler (Südtirol, Italien). Gredleriana, 4, 93–114.

    Schatz, H., Behan-Pelletier, V.M., OConnor, B.M. & Norton, R.A. (2011) Suborder Oribatida van der Hammen, 1968. In: Zhang, Z.Q. (Ed.), Animal biodiversity: An outline of higher level classification and survey of taxonomic richness. Zootaxa, 3148, 141–148.

    Shaldybina, E.S. (1975) Family Galumnidae. In: Ghilarov, M.S. & Krivolutsky, D.A. (Eds.), Identification keys of soil inhabiting mites, Sarcoptiformes. Nauka, Moscow, 347–363. [in Russian]

    Shtanchaeva, U. Ya. (2001) Catalog of oribatid mites of the Caucasus (Acari, Oribatida). Acarina, 9 (2), 177–221.

    Shtanchaeva, U. Ya. (2008) A review of oribatid mites of the family Liacaridae (Acariformes, Oribatida) from the Caucasus. Entomological review, 88 (2), 244–257.
    http://dx.doi.org/10.1134/S0013873808020103

    Shtanchaeva, U. Ya & Subías, L.S. (2010) The catalogue of Caucasian oribatid mites. Nauka, Makhachkala. 276 pp. [in Russian]

    Shtanchaeva, U. Ya., Subías, L.S. & Arillo, A. (2010) New taxa of oribatid mites of the family Liacaridae (Acariformes, Oribatida) from the Caucasus. Entomologica Fennica, 20, 245–248.

    Shtanchaeva, U. Ya. & Subías, L.S. (2012) New species of primitive oribatid mite families Brachychthoniidae and Phthiracaridae (Acariformes, Oribatida) in Caucasus. Entomological Review, 92 (4), 447–458.
    http://dx.doi.org/10.1134/s0013873812040100

    Shtanchaeva, U. Ya. & Subías, L.S. (2012a) A new subgenus and three new species of the oribatid mite families Hermanniellidae, Oribatellidae, and Scheloribatidae (Acariformes, Oribatida) from the Caucasus. Entomological Review, 92 (5), 583–589.
    http://dx.doi.org/10.1134/S0013873812050119

    Subías, L.S. (2004 updated in 2015) Listado sistemático, sinonimico y biogeografico de los oribatidas (Acariformes, Oribatida) del mundo (1758–2002). 2004. Graelsia, 60 (numero extraordinario), 305 pp. Available from: http://www.ucm.es/info/zoo/Artropodos/Catalog.pdf (accessed 1 December 2015)

    Subías, L.S. (2000) Nuevos oribátidos (Acariformes, Oribatida) para la fauna de la Península Ibérica. Graellsia, 56, 21–25.

    Subías, L.S. (2010) Nuevos nombres de oribátidos (Acari: Oribatida). Boletin Real Sociedad Española de Historia Natural. Sección Biológica, 104, 35–39.

    Subías, L.S., Kahwash, M.A.M. & Ruiz, E. (1990) Un nuevo género y tres nuevas especies de Ceratozetoideos del Sur de España (Acari, Oribatida, Ceratozetoidea). Boletín de la Asociación española de Entomología, 14, 105–113.

    Subías, L.S. & Arillo, A. (2000) Ácaros Oribátodes (Acariformes, Oribatida) de la Sierra de Mira (Este de España). Boletín de la Asociación Española de Entomología, 24 (3–4), 85–104.

    Tarba, Z.M. (1974) Vertical distribution of oribatid mites in Caucasian forest soils. Zoologichesky Zhurnal, 53 (11), 1630–1635. [in Russian]

    Tarba, Z.M. (1976) The fauna of oribatid mites of Ritsa-Auadkhara Reserve. Fauna i ekologiya bezpozvonochnikh zhivotnikh. Moskow, 20–24. [in Russian]

    Tarba, Z.M. (1978) The fauna of oribatid mites of Abkhazia. Trudi pedagogicheskikh institutov Gruzinskoi SSR. Seria estestvennikh nauk, 5, 68–81. [in Russian]

    Tarba, Z.M. (1988) The seasonal dynamic of Kolkheti type forest in Abkhazia. Ekologiya mikroartropod lesnikh pochv. Nauka, Moscow, 81–92. [in Russian]

    Tarba, Z.M. (1990) New species of oribatid mites – Heminothrus sp.n. from Abkhazia. Zoologichesky Zhurnal, 89, 142–143. [in Russian]

    Tarba, Z.M. (1993) Population structure of oribatid mites in the vertical landscape zones of Abkhazia. Zoologichesky zhurnal, 72 (8), 22–27.

    Weigmann, G. (2001) Contribution to the knowledge of European Poronota I. Oribatella and Anachipteria. Spixiana, 24 (3), 235–240.

    Weigmann, G. (2006) Hornmilben (Oribatida). Die Tierwelt Deutschlands. 76 Teil. Goecke & Evers, Keltern, 520 pp.

    Weigmann, G., Horak, F., Franke, K. & Christian, A. (2015) Verbreitung und Ökologie der Hornmilben (Oribatida) in Deutschland. Peckiana, 10, 1–171.

    Weigmann, G. & Schatz, H. (2015) Redescription of Coronoquadroppia monstruosa (Hammer, 1979) (Acari, Oribatida, Quadroppiidae) from Java and variability of the species in Europe. Zootaxa, 3926 (3), 329–350.
    http://dx.doi.org/10.11646/zootaxa.3926.3.2

    Willmann, C. (1931) Moosmilben oder Oribatiden (Cryptostigmata). In: Dahl, F. (Ed.), Die Tierweld Deutschlands, 22, 79–200.

    Zazanashvili, N., Sanadiradze, G., Bukhnikashvili A., Kandaurov, A. & Tarkhnishvili, D. (2004) Caucasus. In: Mittermaier, R.A., Hoffmann, P.G. Gil, M., Pilgrim, J., Brooks, T., Mittermaier, C.G., Lamoreux J. & da Fonseca, G.A.B. (Eds.), Hotspots revisited, Earth’s biologically richest and most endangered terrestrial ecoregions, CEMEX/Agrupacion, Sierra Madre, pp. 148–153.