Abstract
Arcanobisium gen. nov. is described for Arcanobisium comasi sp. nov., from a limestone cave in the Valencian Community, Castellón province, Spain. By its unique features, compared to other syarinids, it warrants the creation of a new subfamily, Arcanobisiinae, for its accommodation. Some characters of the Syarinidae are evaluated and the genera Chitrellina Muchmore, 1996, Hadoblothrus Beier, 1952, Microcreagrella Beier, 1961 and Microcreagrina Beier, 1961 are transferred to the subfamily Ideobisiinae from the Chitrellinae. A key to the subfamilies of Syarinidae is given. The relictual status of the new subfamily is discussed.References
Banks, N. (1893) New Chernetidae from the United States. Canadian Entomologist, 25, 64–67.
Beier, M. (1931) Zur Kenntnis der troglobionten Neobisien (Pseudoscorp.). Eos, Madrid, 7, 9–23.
Beier, M. (1932) Pseudoscorpionidea I. Subord. Chthoniinea et Neobisiinea. Tierreich, 57, i–xx, 1–258.
Beier, M. (1952) Über die von L. di Caporiacco aus Apulien beschriebenen Höhlen-Pseudoscorpione. Memorie di Biogeografica Adriática, 2, 103–108.
Beier, M. (1961) Pseudoscorpione von den Azoren und Madeira. Boletim do Museu Municipal do Funchal, 14, 67–74.
Beier, M. (1963) Ordnung Pseudoscorpionidea (Afterskorpione). Bestimmungsbücher zur Bodenfauna Europas. Vol. 1. Akademie-Verlag. Berlin, 313 pp.
Beier, M. (1969) Reliktformen in der Pseudoscorpioniden-Fauna Europas. Memorie della Società Entomologica Italiana, 48, 317–323.
Caporiacco, L. di (1951) Aracnidi cavernicoli Pugliesi. Memorie di Biogeografica Adriática, 2, 95–101.
Chamberlin, J.C. (1925) On a collection of pseudoscorpions from the stomach contents of toads. University of California Publications in Entomology, 3, 327–332.
Chamberlin, J.C. (1930) A synoptic classification of the false scorpions or chela-spinners, with a report on a cosmopolitan collection of the same. Part II. The Diplosphyronida (Arachnida-Chelonethida). Annals and Magazine of Natural History, (10) 5, 1–48, 585–620.
Chamberlin, J.C. (1931) The arachnid order Chelonethida. Stanford University Publications, Biological Sciences, 7(1), 1–284.
Chamberlin, J.C. (1938) A new genus and three new species of false scorpion from Yucatan Caves (Arachnida - Chelonethida). Publications of the Carnegie Institution of Washington, 491, 109–121.
Ćurčić, B.P.M. (1979) The genus Pararoncus Chamberlin 1938 (Pseudoscorpiones, Neobisiidae) in Japan. Glasnik Muzeja Srpske Zemlje, Beograd (B), 34, 169–180.
Ellingsen, E. (1901) Sur une espèce nouvelle d’Ideobisium genre des pseudoscorpions de l’Europe. Bulletin de la Société Zoologique de France, 26, 86–89.
Ellingsen, E. (1908) Two Canadian species of pseudoscorpions. Canadian Entomologist, 40, 163.
Harvey, M.S. (1992) The phylogeny and classification of the Pseudoscorpionida (Chelicerata: Arachnida). Invertebrate Taxonomy, 6, 1373–1435.
Harvey, M.S. (1998) Pseudoscorpion groups with bipolar distributions: a new genus from Tasmania related to the Holarctic Syarinus (Arachnida, Pseudoscorpiones, Syarinidae). Journal of Arachnology, 26, 429–441.
Harvey, M.S. (2009) Pseudoscorpions of the World, version 1.2. Western Australian Museum, Perth. Available from: http://www.museum.wa.gov.au/arachnids/pseudoscorpions.
Harvey, M.S. & Edward, K.L. (2007) A review of the pseudoscorpion genus Ideoblothrus (Pseudoscorpiones, Syarinidae) from western and northern Australia. Journal of Natural History, 41, 445–472.
Harvey, M.S. & Volschenk, E.S. (2007) The systematics of the Gondwanan pseudoscorpion family Hyidae (Pseudoscorpiones: Neobisioidea): new data and a revised phylogenetic hypothesis. Invertebrate Systematics, 21, 365–406.
Hong, Y. & Kim, T.H. (1996) On a species in the genus Pararoncus (Pseudoscorpionida: Neobisiidae) from Korea. Korean Journal of Soil Zoology, 1, 134–139.
Inzaghi, S. (1983) Pseudoblothrus regalini n. sp., da grotte della Provincia di Bergamo (Italia sett.) (Pseudoscorpiones Syarinidae). Atti della Società Italiana di Scienze Naturali, e del Museo Civile di Storia Naturale, Milano, 124, 38–48.
Judson, M.L.I. (2000) Electrobisium acutum Cockerell, a cheiridiid pseudoscorpion from Burmese amber, with remarks on the validity of the Cheiridioidea (Arachnida, Chelonethi). Bulletin of the Natural History Museum, Geology, 56, 79–83.
Judson, M.L.I. (2007) A new and endangered pseudoscorpion of the genus Lagynochthonius (Arachnida, Chelonethi, Chthoniidae) from a cave in Vietnam, with notes on chelal morphology and the composition of the Tyrannochthoniini. Zootaxa, 1627, 53–68.
Mahnert, V. (1976) Zur Kenntnis der Gattungen Acanthocreagris und Roncocreagris (Arachnida, Pseudoscorpiones, Neobisiidae). Revue Suisse de Zoologie, 83, 193–214.
Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719–810.
Mahnert, V. (1980) Pseudoskorpione (Arachnida) aus Höhlen Italiens, mit Bemerkungen zur Gattung Pseudoblothrus. Grotte d'Italia, (4a) 8, 21–38.
Mahnert, V. (1985) Weitere Pseudoskorpione (Arachnida) aus dem zentralen Amazonasgebiet (Brasilien). Amazoniana, 9, 215–241.
Mahnert, V. (1990) Deux nouvelles espèces du genre Pseudoblothrus Beier, 1931 (Pseudoscorpiones, Syarinidae) des Açores (Portugal). Vieraea, Tenerife, 18, 167–170.
Mahnert, V. (1993) Pseudoskorpione (Arachnida: Pseudoscorpiones) von Inseln des Mittelmeers und des Atlantiks (Balearen, Kanarische Inseln, Madeira, Ascension), mit vorwiegend subterraner Lebensweise. Revue suisse de Zoologie, 100, 971–992.
Mahnert, V. (2008) Pseudoscorpiones. In: Borges, P.A.V., Abreu, C., Aguiar, A.M.F., Carvalho, P. Jardim, R., Melo, I., Oliveira, P. Sergio, C. Serrano, A.R.M. & Vieira, P. (Eds.), A list of the terrestrial fungi, flora and fauna of Madeira and Selvagens archipelago. Direcção Regional do Ambiente da Madeira and Universidade dos Açores, Funchal and Angra do Heroismo, p. 279.
Malcolm, D.R. & Chamberlin, J.C. (1960) The pseudoscorpion genus Chitrella (Chelonethida, Syarinidae). American Museum Novitates, 1989, 1–19.
Muchmore, W.B. (1973) The genus Chitrella in America (Pseudoscorpionida, Syarinidae). Journal of the New York Entomological Society, 81, 183–192.
Muchmore, W.B. (1979) Pseudoscorpions from Florida and the Caribbean area. 7. Floridian diplosphyronids from Florida and the Caribbean area. Florida Entomologist, 62, 193–213.
Muchmore, W.B. (1982) The genera Ideobisium and Ideoblothrus, with remarks on the family Syarinidae (Pseudoscorpionida). Journal of Arachnology, 10, 193–221.
Muchmore, W.B. (1990) Pseudoscorpionida. In: Dindal, D.L. (Ed.), Soil biology guide. John Wiley and Sons, New York, pp. 503–527.
Muchmore, W.B. (1992) Cavernicolous pseudoscorpions from Texas and New Mexico (Arachnida: Pseudoscorpionida). Texas Memorial Museum, Speleological Monographs, 3, 127–153.
Muchmore, W.B. (1996) A remarkable new genus and species of Pseudoscorpionida (Syarinidae) from a cave in Arizona. Southwestern Naturalist 41, 145–148.
Murienne, J., Harvey, M.S. & Giribert, G. (2008) First molecular phylogeny of the major clades of Pseudoscorpiones (Arthropoda: Chelicerata). Molecular Phylogenetics and Evolution, 49, 170–184.
Ortuño, V.M., Sendra, A., Montagud, S. & Teruel, S. (2005) Systématique et biologie d’une espèce paléoendémique hypogée de la péninsule Ibérique: Ildobates neboti Español, 1966 (Coleoptera: Carabidae: Dryptinae). Annales de la Société Entomologique de France (Nouvelle Série), 40, 459–475.
Ortuño, V.M., Sendra, A., Bellés, X., Montagud, S. & Teruel, S. (2006) Ildobates neboti Español, 1966 (Coleoptera: Carabidade: Zuphiini): un icono de la bioespeleología ibérica. Boletín Sociedad Entomológica Aragonesa, 38, 379−382.
Redikorzev, V. (1918) Pseudoscorpions nouveaux. I. Ezhegodnik Zoologicheskago Muzeya, 22, 91–101.
Schawaller, W. (1987) Erstnachweis der Familie Syarinidae in Deutschland: ein Reliktvorkommen von Syarinus strandi im Oberen Donautal (Arachnida: Pseudoscorpiones). Jahreshefte, Gesellschaft für Naturkunde in Württemberg, 142, 287–292.
Sendra, A. (2003) Distribución y colonización de los Campodeidos cavernícolas en la Península Ibérica e Islas Baleares. Boletín de la Sociedad Española de Espeleología y Ciencias del Karst, 4, 12–20.
Sendra, A. (2006) Arthropoda, Entognatha, Diplura. In: Domingo J., Montagud, S., S. & Sendra, A. (Eds.) 2006. Invertebrados endémicos de la Comunitat Valenciana. Generalitat Valenciana, Conselleria de Territori i Habitatge. Generalitat Valenciana, pp. 124–130.
Sendra, A., Ortuño, V.M., Moreno, A., Montagud, S. & Teruel, S. (2006) Gollumjapyx smeagol gen. n., sp. n., an enigmatic hypogean japygid (Diplura: Japygidae) from the eastern Iberian Peninsula. Zootaxa, 1372, 35–52.
Simon, E. (1883) Matériaux pour servir à la faune arachnologique des îles de l’Océan Atlantique (Açores, Madère, Salvages, Canaries, Cap Vert, Sainte-Hélène et Bermudes). Annales de la Société Entomologique de France, (6) 3, 259–314.
Vachon, M. (1952) Remarques préliminaires sur 1’anatomie et la biologie de deux Pseudoscorpions très rares de la faune française: Pseudoblothrus peyerimhoffi (E. S .) et Apocheiridium ferum (E. S.). Bulletin du Muséum National d'Histoire Naturelle, Paris (2) 24, 536–539.
Vachon, M. (1954) Remarques morphologiques et anatomiques sur les Pseudoscorpions (Arachnides) appartenant au genre Pseudoblothrus (Beier) (Fam. Syarinidae J.C.C.) (à propos de la description de P. strinatii n. sp., des cavernes de Suisse). Bulletin du Muséum National d'Histoire Naturelle, Paris, (2) 26, 212–219.
Vachon, M. (1969) Remarques sur la famille des Syarinidae J.C. Chamberlin (Arachnides, Pseudoscorpions) à propos de la description d’une nouvelle espèce: Pseudoblothrus thiebaudi habitant des cavernes de Suisse. Revue Suisse de Zoologie, 76, 387–396.
Zaragoza, J.A. (1990) Escorpiones, pseudoescorpiones y solífugos de la provincia de Alicante. Ayudas a la investigación 1986-1987. Instituto de Cultura Juan Gil-Albert. Diputación Provincial de Alicante, Alicante, vol. 1, pp. 45–70.
Zaragoza, J.A. (2004) Pseudoscorpiones. In: Barrientos, J.A. (Ed.), Curso práctico de entomología. Universitat Autònoma de Barcelona, Barcelona, pp. 177–187.
Zaragoza, J.A. (2007) Catálogo de los Pseudoescorpiones de la Península Ibérica e Islas Baleares (Arachnida: Pseudoscorpiones). Revista Ibérica de Aracnología, 13, 3–91.
Zaragoza, J.A. (2008) On the status of the subspecies of Roncocreagris galeonuda (Pseudoscorpiones: Neobisiidae): importance of the chelal microsetae pattern. Remarks on the genus Roncocreagris Mahnert. Revista Ibérica de Aracnología, 15, 35–46.