Zootaxa

**Revision of the Lethrus (Heteroplistodus) crenulatus-species group (Coleoptera: Geotrupidae: Lethrinae). Part 1: species similar to L. tschitscherini Semenov, 1894**

Wed, 17 Jun 2026 00:00:00 +1200

The Lethrus crenulatus-species group is discussed. Lethrus (Heteroplistodus) ikabaki A. Shapovalov, Kolov, Hillert & Král, sp. nov. is described from China (Xinjiang Province), L. (H.) heptapotamicus A. Shapovalov, Bagaturov, Hillert & Král, sp. nov. and L. (H.) tengri A. Shapovalov, Hillert & Král, sp. nov. are described from southeastern Kazakhstan. A key to the species distributed in the Ili Intermontane Basin is provided. Relevant characteristics are illustrated with photographs.

Praying Mantises (Insecta: Mantodea) of the Eastern Himalayas: A Systematic Review of Diversity and Range Extensions from Arunachal Pradesh

P.M. SURESHAN, C. BINOY, S.D. GURUMAYUM — Wed, 17 Jun 2026 00:00:00 +1200

Praying Mantises (Insecta: Mantodea) are an important yet understudied component of predatory arthropod assemblages in the Eastern Himalaya. We provide a systematic account of the mantid fauna of Arunachal Pradesh, India, based on the study of mantis specimens collected from different parts of the state such as Tawang district, Namdapha Tiger Reserve, Kane Wildlife Sanctuary, Yordi-Rabe Subse Wildlife Sanctuary, and Upper Siang District supplemented by literature review. A total of 29 species across 18 genera and five families are documented. Field collections yielded 74 specimens, confirming known records and reporting seven new distributional records for the state: Amantis subirina, Acromantis lobofemorata, Anaxarcha graminea, Astyliasula inermis, Ephestiasula rogenhoferi, Hierodula tenuidentata, and Tenodera aridifolia. The assemblage reflects the biogeographical complexity of Arunachal Pradesh, which hosts a mix of Oriental and Palearctic elements, providing baseline data for biodiversity assessment and conservation.

Spatulosia Toulgoёt, 1965, a confirmed Malagasy genus, new for continental Tropical Africa, with description of four new species (Lepidoptera: Erebidae: Arctiinae: Lithosiini)

ANTONIO DURANTE — Wed, 17 Jun 2026 00:00:00 +1200

The genus Spatulosia Toulgoёt, 1965 is re-described owing to the inclusion of three already-described species (Spatulosia africana (Hampson, 1900) comb. nov.; Spatulosia coronaria (Kühne, 2007) comb. nov.; Spatulosia gemina (Hampson, 1914) comb. nov.) and the discovery of four new species (Spatulosia meridionalis sp. nov.; Spatulosia occidentalis sp. nov.; Spatulosia pallida sp. nov.; Spatulosia scutulata sp. nov.). The genus is compared with similar or related genera and is validated with reference to probable synapomorphies. Its original Malagasy range is enlarged to include the whole of Sub-Saharan Africa.

**A taxonomic study of the genus Isturgia from China (Lepidoptera, Geometridae, Ennominae)**

XIAOYE MI, RUI CHENG, HONGXIANG HAN — Wed, 17 Jun 2026 00:00:00 +1200

The genus Isturgia Hübner, 1823 in China is reviewed. Ten species are recognized, of which, Isturgia bisinuata sp. nov. from Xizang is described as new to science. Two new combinations are established (Isturgia anostilpna (Wehrli, 1938) comb. nov., and Isturgia isogrammica (Wehrli, 1937) comb. nov.) and Isturgia murinaria is reported as a new record for China. Diagnoses for all species are provided, and illustrations of external features and genitalia are presented.

**Review of the genus Silusa Erichson (Coleoptera: Staphylinidae: Aleocharinae) from Japan, with notes on species described from India**

TAKUTO HASHIZUME, MUNETOSHI MARUYAMA — Wed, 17 Jun 2026 00:00:00 +1200

The genus Silusa Erichson, 1837 in Japan is revised based on the examination of type material and additional specimens. Four species are recognized as valid members of the genus, excluding species with highly doubtful generic assignment: Silusa (Silusa) biimpressa Bernhauer, 1907 = Silusa chinensis Pace, 1998, syn. nov., Silusa (Silusa) lanuginosa Sharp, 1888, Silusa immutata sp. nov., and Silusa variabilis sp. nov. Both male and female genital structures of the Japanese species are redescribed in detail, allowing reliable identification for the first time. To clarify the status of South Asian taxa, four species described from India by Cameron (1939) as Silusa, are here illustrated and briefly diagnosed with additional descriptions: Silusa (Silusa) infuscata Cameron, 1939, Silusa indica Cameron, 1939, Coenonica distincta (Cameron, 1939), comb. nov. = Coenonica arcusifera Pace, 1998, syn. nov., and “Silusa” nigra Cameron, 1939 (transferred to Oxypodini Thomson). This study resolves long-standing taxonomic confusion in the Japanese Silusa species and provides an updated diagnostic framework for East and South Asian species of the genus.

**Review of the Palaearctic species of Dicopomorpha Ogloblin, 1955 (Hymenoptera: Mymaridae)**

FILIP PAWLUK, DAWN PAINTER — Wed, 17 Jun 2026 00:00:00 +1200

The fairyfly genus Dicopomorpha Ogloblin (Hymenoptera: Mymaridae) is recorded for the first time from Europe. The female of D. liaoningensis (Lou, Cao & Lou), originally described from Northeastern China, is redescribed based on specimens from New Zealand, Poland and United Kingdom, and its previously unknown male is described and illustrated. Litus longitus Aishan is transferred to Dicopomorpha as D. longita (Aishan), comb. nov. A key to females of the three Palaearctic species of Dicopomorpha is presented.

A new species of the family Saturniidae from Hainan Island (Lepidoptera: Saturniidae)

QI-LE HE — Wed, 17 Jun 2026 00:00:00 +1200

In March 2025, unknown Saturniidae larvae were found on Jianfengling Mountain, western Hainan Island, and reared to adulthood, revealing a complete life history. Morphological comparisons and COI barcode‑based phylogenetic analyses supported the new taxon as an independent lineage. The new species is characterized by unique male genitalia (vesica with a deeply embedded, never fully everted cornutus), and distinctive larval characters (semi‑fused dorsal scoli on A8, extensive black coloration). Herein described as Cricula qiongya sp. nov., it is currently known only from a restricted area on Jianfengling Mountain.

**A new Phyllogomphoides Belle from Ecuador, with description of male, female and larva (Odonata: Gomphidae)**

GÜNTHER FLECK, WILLIAM A. HABER — Wed, 17 Jun 2026 00:00:00 +1200

Phyllogomphoides biguttatus sp. nov., from western Ecuador, is described based on reared adults of both sexes and larvae collected in slow parts of small forest streams in hilly landscapes. This species belongs to the semicircularis-group of species and is closely related to P. semicircularis (Selys Longchamps) and P. insignatus Donnelly. Phyllogomphoides insignatus probably represents the adelphotaxon of the new species. A differential diagnosis is given to distinguish these closely related species. The new species can be separated from all known Phyllogomphoides Belle by its unique thoracic color pattern with pleural and tergal parts of pterothorax entirely brown to dark brown except for a pair of large yellow drop-shaped antehumeral markings.

**A new species of the genus Termitodiellus Nakane, 1961 (Coleoptera: Scarabaeidae: Aphodiinae) from Vietnam**

ŁUKASZ MINKINA, ADAM BYK, MARCIN JAN KAMIŃSKI — Wed, 17 Jun 2026 00:00:00 +1200

A new Termitodiellus Nakane, 1961 species from Vietnam is described and illustrated: Termitodiellus bielawskipisars-kiorum Minkina, Byk & Kamiński, sp. nov. New distributional records for Termitodiellus luzonensis (Howden, 1995) are provided. The taxonomic complexities of Termitodiellus are briefly discussed in the light of related genera and an updated key for species identification is presented. Finally, a map of the distribution of all currently known Termitodiellus species is provided.