Zootaxa

Unveiling the secrets of South American Darwin wasps, part I: a comprehensive checklist of the Chilean Ichneumonidae (Hymenoptera: Ichneumonoidea)

RODRIGO O. ARAUJO — 2025-05-01

For the first time, a checklist of the species of Chilean Ichneumonidae (Hymenoptera: Ichneumonoidea) is provided, resulting from 10 years of dedicated efforts to visit collections and gather scientific articles on Chilean species. Geographical data for all species were compiled from all bibliographic references that mention Chilean species up to the present. A total of 306 species belonging to 141 genera and 23 subfamilies are registered in Chile, of which 209 species, 30 genera, and one subfamily are endemic. Tycherus Förster is the richest genus in Chile, with 35 species, followed by Alophophion Cushman (22 species), Mesochorus Gravenhorst (20 species), Lepidura Townes (13 species), and Trachysphyrus Haliday (13 species). These five genera have undergone taxonomic revisions, suggesting that the total number of Chilean species is likely underestimated and is expected to significantly increase through future revisions, particularly focusing on poorly studied genera. This comprehensive checklist not only consolidates existing data but also serves as a crucial resource for future taxonomic research, conservation efforts, and effective management of the diverse Darwin wasps in Chile. A comprehensive understanding of the distribution, abundance, and endemism of Ichneumonidae is essential for accurately assessing the ecological dynamics of the country and implementing targeted conservation strategies.

Further discussion of relationships within Australasian Neopilionidae (Opiliones: Phalangioidea), with description of two new species and eight new genera

CHRISTOPHER K. TAYLOR — 2025-05-01

Two new species of Enantiobuninae (Opiliones: Neopilionidae) are described from New Zealand, Accensus brevicrus new genus and species and Maikukunui tokerau new genus and species, and included in an updated morphological phylogenetic analysis of Australasian Enantiobuninae. The genera Megalopsalis and Forsteropsalis as previously recognised are not supported as monophyletic. Classification of the Enantiobuninae is revised in view of morphological data and comparison with prior molecular phylogenetic studies. Megalopsalis is divided between three morphologically diagnostic subgenera, including M. (Intutoportula) new subgenus and M. (Spinicruroides) new subgenus. Megalopsalis nigricans is transferred to Spinicrurellum new genus and M. triascuta to Triascutum new genus. Forsteropsalis is also subdivided with F. grimmetti assigned to Pakaka new genus, F. inconstans to Shelob new genus, F. pureora to Puwere new genus and F. photophaga and related taxa to Ungoliant new genus. Dasylobus australis is provisionally recognised as Forsteropsalis australis new combination. A key is provided to New Zealand species of Palpatores.

Still undescribed bees in Europe—description of nine new European osmiine bee species (Hymenoptera, Megachilidae)

ANDREAS MÜLLER — 2025-05-01

Bees of the tribe Osmiini (Megachilidae) are represented in Europe by about 250 described species. In this publication, nine additional European osmiine bee species are newly described and diagnosed: Heriades (Michenerella) clypeonitida sp. nov. from Spain, southeastern Europe and western Turkey, Hoplitis (Anthocopa) nevadensis sp. nov. and Hoplitis (Anthocopa) spinidorsa sp. nov. from the Sierra Nevada in southern Spain, Hoplitis (Hoplitis) cretensis sp. nov. from Crete, Hoplitis (Micreriades) andalusiae sp. nov. from southwesternmost Andalusia, Hoplitis (Micreriades) hispaniae sp. nov. and Hoplitis (Micreriades) fuscospina sp. nov. from central and southern Spain, Osmia (Helicosmia) inflatipes sp. nov. from Greece, and Osmia (Helicosmia) sardoa sp. nov. from Sardinia. For Hoplitis convolvuli Müller, 2022 (not Osmia papaveris convolvuli Ducke, 1899 = Osmia perezi Ferton, 1895), a replacement name is given: Hoplitis (Anthocopa) trabutiana nom. nov..

Reorganization of New World Helopini (Coleoptera: Tenebrionidae: Tenebrioninae)

ROLF L. AALBU — 2025-05-01

The New World Helopini are reorganized following the examination of the holotype from the neglected taxon Dinomus Brême, 1842 (type species Dinomus perforatus Brême, 1842, by monotypy). It has been known for some time that the New World taxa, mostly treated under the genus Helops Fabricius, 1775, are unrelated to the Palearctic type species of the genus. Our investigations lead us to recognize two genera: Dinomus which contains most of the known Nearctic and Neotropical helopine fauna and Nalassus Mulsant, 1854, a primarily Palearctic genus with several Nearctic species. The following genera are synonymized under the genus Dinomus Brême, 1842: Doyenellus, Nabozhenko & Steiner, 2021 syn. nov.; Neohelops, Dajoz, 2001 syn. nov.; Nautes Pascoe, 1866 syn. nov.; and Tarpela Bates, 1870 syn. nov. Other North American genera previously synonymized under Helops Fabricius, 1775 are here moved into synonymy with Dinomus: Stenotrichus LeConte, 1862 syn. nov.; Biomorphus Motschulsky, 1872 syn. nov.; Coscinoptilix Allard, 1876 syn. nov.; and Lamperos Allard, 1876, previously synonymized under Tarpela Bates, 1870, syn. nov. All species in the previous synonymized genera are transferred to Dinomus Brême. All other current New World species identified as belonging to the genus “Helops” are transferred to the genus Dinomus Brême. Helops perforatus Horn, 1880 from Texas is renamed Dinomus horni Aalbu & Johnston nom. nov. A table is presented for all specific name changes in the New World Helopini.

**Four new species of the genus Hexarhopalus Fairmaire, 1891 (Coleoptera, Tenebrionidae, Stenochiinae) from Vietnam and India**

LUBOŠ PURCHART — 2025-05-01

Three new species of the genus Hexarhopalus—H. (Hexarhopalus) kaminskii sp. nov., H. (Hexarhopalus) ruzzieri sp. nov. and H. (Leprocaulus) kontumensis sp. nov. from Vietnam and one new species—H. (Leprocaulus) arunachalpradeshensis sp. nov. from India are described. First additional specimens of H. masumotoi Bečvář & Purchart, 2008, H. lilligi Bečvář & Purchart, 2008 and H. sinjaevi Bečvář & Purchart, 2008 are reported since their description based on a single or two type specimens, respectively. H. lilligi is reported as new for Laos. A lectotype is designated for Derosphaerus binhanus Pic, 1922. The latter species is redescribed and transferred to the genus Hexarhopalus as H. binhanus (Pic, 1922) comb. nov. and reported as new for Thailand and Laos. An updated checklist of all known Hexarhopalus species is provided.

**Female association of seven species of the genus Amphinemura Ris, 1902 (Nemouridae: Amphinemurinae) in China based on morphological and molecular data**

BINGLI WANG — 2025-05-01

Females of seven known species of nemourid stoneflies, Amphinemura claviloba (Wu, 1973), A. dabanshana Li, Du & Yang, 2017, A. hastata (Wu, 1973), A. multispina (Wu, 1973), A. ningxiana Li & Yang, 2011, A. pediformis Li & Yang, 2008 and A. qiliana Li, Teslenko & Yang, 2020, are described in this paper. DNA barcoding analysis confirmed the association between males and females, thus validating the morphological taxonomic conclusions.

**A new species of Aspidomorphus (Serpentes: Elapidae) from Papua New Guinea**

FRED KRAUS — 2025-05-01

Papuan snakes of the genus Aspidomorphus have long been taxonomically problematic, but nomenclature within this group has been stable at three recognized species since 1967. Previous genetic work suggested the presence of a number of cryptic but currently unrecognized candidate species in the genus. In 2004, I obtained a series of Aspidomorphus from Sudest Island, Milne Bay Province, Papua New Guinea that were clearly different from currently recognized species, and I describe that species here as A. dimorphus sp. nov. This species exhibits a striking bimodality in color pattern in adults, here termed the “orange morph” and “dark morph”. This dimorphism is not related to sex, and both forms have earlier been shown to be genetically identical. Description of A. dimorphus sp. nov. begins the process of bringing taxonomic clarity to a genus long held in the grip of nomenclatural stasis. It furthermore highlights again the importance of Sudest Island for biodiversity endemism.

Holotrichia burmeisteri Brenske, 1892 and allies (Coleoptera: Scarabaeidae: Melolonthinae: Rhizotrogini): a tale of four genera and four countries

PETER G. ALLSOPP — 2025-05-01

The Palauan species Holotrichia burmeisteri Brenske, 1892 is shown to be a member of the Rhizotrogini and, hence, incorrectly placed in the eastern Australian genus Metatrogus Britton, 1978 (Melolonthini). Comparison with another species described from Palau shows that H. burmeisteri is a senior synonym of Lepidiota carolinensis Arrow, 1939 (new synonymy). That species and two other closely related species also described from Palau, L. dybasi Gordon, 1979 and L. furtiva Gordon, 1979, are transferred to Cheberdord Allsopp, new genus. The presumed occurrence of C. burmeisteri in the Philippines and Australia is considered extremely doubtful, and the record from Guam is based on one individual transported in luggage. Lectotypes are designated for Holotrichia burmeisteri Brenske, 1892 and Lepidiota carolinensis Arrow, 1939.

**Taxonomy of the genus Phloeophagus Schoenherr (Curculionidae: Cossoninae) from Japan, with description of a new species**

SHÛGO INOUE — 2025-05-01

The Japanese species of the genus Phloeophagus Schoenherr, 1838 are revised taxonomically. Two species, P. japonicus Inoue, sp. nov. (Hokkaido, Honshu, and Kyushu) and P. orientalis Osella, 1974, are recognized. Both species are described or redescribed.

Moema humaita, a new species of annual fish (Cyprinodontiformes: Rivulidae) from the middle rio Madeira, Amazon basin, Brazil

DALTON TAVARES BRESSANE NIELSEN — 2025-05-01

Moema humaita, new species, is described from a seasonal pool in the rio Madeira drainage, Amazon River basin, Brazil. The new species is assigned to the Moema staecki species group based on the color pattern in males, composed of dots following interrupted lines arranged horizontally. Moema humaita differs from all known congeners by its unique color pattern in males, presenting body sides light grey, with three horizontal lines, composed by small red dots lying in the center of each scale, alternating with three bright yellow lines, and additionally by presenting caudal fin with 3–4 long filamentous rays and presenting an anal fin long, with its tip pointed, reaching vertical through near the end of the caudal fin, orange in color. The occurrence of an undescribed species of the genus Moema in Guyana is reported based on a photograph of an unpreserved specimen.

**Comment on “Two new species of Staphylus (Scantilla) Godman and Salvin, 1896 (Lepidoptera, Hesperiidae, Pyrginae, Carcharodini) from Colombia and Mexico” by Lemes et al. (2025)**

JOSÉ RICARDO ASSMANN LEMES — 2025-05-01

**New replacement names for Schaufussia Fresneda et al., 2024, and Dupreia Fresneda et al., 2024 (Coleoptera, Leiodidae, Cholevinae, Leptodirini)**

JAVIER FRESNEDA — 2025-05-01