Zootaxa

**Reassessment of Incasarus (Hymenoptera: Andrenidae: Protandrenini) with the description of two new species (Hymenoptera: Andrenidae)**

NORA ROMERO — 2026-06-09

A revised classification is presented for a group of Andean bees from Argentina, Bolivia, and Peru previously assigned to Liphanthus Reed (Andrenidae: Panurginae: Protandrenini) based on a phylogenetic analysis of 108 morphological characters. The results indicate that these species form a well-supported clade with the previously monotypic genus Incasarus Gonzalez, Rasmussen & Engel, rendering Liphanthus, as currently delimited, paraphyletic. These species share several distinctive morphological characters, including dense to crowded punctation on the head and mesoscutum, strongly sculptured tegulae, and unique genitalic features not observed in Liphanthus. Accordingly, we transfer the following species to a redefined Incasarus: I. abotorabi (Mir Sharifi & Packer, 2019) comb. nov.; I. aeneiventris (Friese, 1908) comb. nov. treated here as the senior synonym of L. fritzi Mir Sharifi & Packer, 2019 syn. nov.; I. amblayensis (Mir Sharifi & Packer, 2019) comb. nov.; I. ancashensis (Mir Sharifi & Packer, 2019) comb. nov.; I. cochabambensis (Mir Sharifi & Packer, 2019) comb. nov.; and I. yrigoyeni (Packer, 2019) comb. nov. Additionally, we describe two new species: I. arequipensis Romero & Packer, sp. nov. from Peru, and I. ivari Romero & Packer, sp. nov. from Bolivia, producing a generic richness of nine species. The phylogenetic analysis suggests three distinct lineages within Incasarus. This study enhances our understanding of the diversity of Protandrenini in the Andes and highlights the continued value of museum material and new field sampling for the discovery of diversity within poorly understood lineages.

Three new species of Cyprettadopsini of subfamily Cypridopsinae (Crustacea: Ostracoda) from Northeast Thailand with discussion on characters in the tribe

SUKONTHIP SAVATENALINTON — 2026-06-09

A survey of the ostracod fauna in the Chi River Basin in three provinces (Maha Sarakham, Kalasin and Roi Et) of northeast Thailand uncovered three new species of the tribe Cyprettadopsini of the Cypridopsinae: Cyprettadopsis hanjavanitae sp. nov., C. kalasinensis sp. nov. and Pseudocypretta sangpradubae sp. nov. Apart from its different soft part morphology, P. sangpradubae sp. nov. can be obviously distinguished from other Pseudocypretta by the valve ornamentation with strong pits and densely arranged long setae. The two new Cyprettadopsis species can be primarily discriminated from the type species, Cyprettadopsis sutura Savatenalinton, 2020, by their subtriangular and subovate carapace shape in, respectively, lateral and dorsal views. Cyprettadopsis hanjavanitae sp. nov. mainly differs from C. kalasinensis sp. nov. by soft parts aspects, especially the caudal ramus. The discovery of these three new species, together with a literature review, provides additional diagnostic features of the Cyprettadopsini. Most of them include the dorsal subapical seta on the basal segment of the maxilla palp, the g seta on the third thoracopod, the pseudosegmentation on the natatory setae of the second antenna, and the color pattern on the valve surface. In addition, setae with thick bases, which are found on the valve surfaces of all species of Cyprettadopsini, have so far been recognized only in the Cypridopsinae. Although the occurrence of marginal septa is an interspecific variable feature in Pseudocypretta and Cyprettadopsis, the modes of appearance are specific to their genera. Herein, the distinguishing features and a key to the genera of the Cyprettadopsini are updated, and Mixtadopsis franki (Ferreira et al., 2023) nov. comb. is introduced. Pseudocypretta sangpradubae sp. nov. is the second Pseudocypretta described from Southeast Asia, and the first one from Thailand. Currently, the number of Cypridopsinae species in Thailand is 15, belonging to six (of a total of seven) tribes, indicating the high diversity at the tribe level of this subfamily in Thailand.

Montana tomorri and Montana dani (Orthoptera: Tettigoniidae), two new species of bush-crickets from the mountains of south Albania

GELLÉRT PUSKÁS — 2026-06-09

Two new taxa of Platycleidini have been discovered in southern Albania, with one of them extending its range to the Greek border. In this paper, we analyse their morphological features, the characteristics of their acoustic signals, and phylogenetic relationships. These taxa clearly belong to the cluster of species comprising the four genera Montana, Amedegnatiana, Parnassiana, and Metrioptera s. str. The results of this study suggest their placement within the genus Montana, and we describe two new species: Montana tomorri sp. nov. and Montana dani sp. nov. Our findings highlight the urgent need for a reassessment of the tribe Platycleidini using integrative approaches that incorporate molecular and bioacoustic data, in order to better understand monophyletic groups and evaluate the consistency of current generic classifications.

New species and new records of tiger beetles of the subgenus Leptinomera Rivalier, 1961 and remarks on the subgenus Matalinia Wiesner & Lien, 2023 of the genus Cylindera Westwood, 1831 (Coleoptera, Cicindelidae)

ANDREY V. MATALIN — 2026-06-09

New records of some species of tiger beetles of the subgenus Leptinomera Rivalier, 1961 from Southeast Asia are presented and discussed. Cylindera (Leptinomera) catoptroides (W. Horn, 1892) is recorded for the first time from Thailand (Nakhon Si Thammarat and Yala Provinces), and Cylindera (Leptinomera) macrodonta Cassola et Probst, 1995 is recorded for the first time from Malaysia (Sabah Province). Three species are described as new: Cylindera (Leptinomera) mosolovi sp. nov.—from Malayan Province Pahang, Cylindera (Leptinomera) pseudocatoptroides sp. nov.—from Thai Province Yala, and Cylindera (Leptinomera) prosvirovi sp. nov.—from Indonesian Provinces Acen and North Sumatra. An identification key as well as the distribution maps of the species of subgenus Leptinomera are provided. Cylindera kualatahanensis Matalin & Cassola, 2000 is transferred from the subgenus Leptinomera to the subgenus Matalinia Wiesner & Lien, 2023.

**The pollen wasps (Hymenoptera: Vespidae: Masarinae) in Central Asia. Contribution II. Genus Celonites Latreille, 1802**

ALEXANDER V. FATERYGA — 2026-06-09

The present contribution continues the revision of the subfamily Masarinae in Central Asia and deals with the genus Celonites Latreille, 1802. This genus includes 14 species: C. biinterruptus Kostylev, 1940, C. crenulatus Morawitz, 1888, C. foveolatus Kostylev, 1935, C. hystrix Kostylev, 1940, C. kozlovi Kostylev, 1935, C. kurzenkoi Mauss, Fateryga & Carpenter, 2026, C. maussi Fateryga, sp. nov., C. mediasiaticus Fateryga, sp. nov., C. modestus Kostylev, 1935, C. montanus Mocsáry, 1906, C. octoannulatus (Kuznetzov, 1923), C. osseus Morawitz, 1888, C. sibiricus Gusenleitner, 2007, and C. tristiculus Kostylev, 1935. Celonites hystrix is reported from Uzbekistan for the first time; C. octoannulatus is reported from Kyrgyzstan for the first time but the identification of the specimens from this country requires further confirmation. An identification key and maps detailing the geographic distribution of all species in Central Asia are presented. Male genitalia and sternum VII+VIII are illustrated for all species (except C. modestus, for which the male is unknown). Taking into account the previously revised genera Masaris Fabricius, 1793 and Quartinia André, 1884, there are three genera and 28 species of pollen wasps in Central Asia; 25 of them are endemic to this region. The largest number of species of pollen wasps among Central Asian countries is known from Kazakhstan (12); there are 11 species in Uzbekistan, 10 species in Tajikistan, nine species in Turkmenistan, five species in Kyrgyzstan, three species in Mongolia, and just one species known from China (Gansu). The biology of the Central Asian species of Masarinae is poorly investigated; nesting is completely unknown for any species occurring there. Any data on trophic relationships are known for 16 species, while just five of them are with rather well-studied flower-visiting behavior and forage preferences.

**DNA barcoding reveals a new species of deepwater catshark of the genus Apristurus (Chondrichthyes: Pentanchidae) from India, in the Southeastern Arabian Sea**

SWETA BEURA — 2026-06-09

This article provides a description of a new species of a catshark of the family Pentanchidae, Apristurus drona sp. nov. based on 4 specimens, which include two males and two females ranging from 439–473 mm TL, collected off the Kollam slope, Southwest coast of India, at a depth of 400–650 m. This new species is placed in the Apristurus brunneus subgroup among the three groups divided within the genus Apristurus and is characterized by the following features: slender body, tapering posteriorly; nostril length is smaller than internarial width; pre-outer nostril length shorter than interorbital space; pre-outer nostril length 3.7(3.5–5.2) % TL; pre-oral length shorter than the mouth width, 33.6(31.7–34.0)% head length and 8.1(7.1–8.1)% TL; nostril-mouth space shorter than nostril length, about 0.4 (0.3–0.4) times; upper labial furrows longer than the lowers furrows; internarial width slightly greater than or equal to orbit length, about 1.09(1.0–1.11) times; 5th gill slit height smallest, about 46.8(46.8–62.5)% eye length; Inter-dorsal space 9.3(8.8–10.5) % TL; first dorsal fin smaller than the second dorsal fin; first dorsal fin insertion is opposite to anal fin origin; anal fin base length is greater than pectoral-pelvic space and pelvic-anal space; pectoral to pelvic space greater than pelvic to anal space; spiral valve turns 11–12; total vertebrae 105–115. Apristurus drona sp. nov. is also distinguished from its congeners based on DNA sequence divergence of the COI gene. This new species has a distribution along the Kollam slope, Arabian Sea, whereas its three closest congeners, based on a molecular level, are from distant locations: A. nayakai from Southwestern Pacific Ocean, A. macrorhynchus from Northwest Pacific and A. exsanguis from New Zealand.

**Redescription of Cubanomysis manuelortizi Hendrickx & Hernández-Payán, 2023 (Crustacea, Mysida, Mysidae) based on a new record from Southeast Gulf of California, Mexico**

JOSÉ CARLOS HERNÁNDEZ-PAYÁN — 2026-06-09

Additional, abundant material of a recently described species of Mysida, Cubanomysis manuelortizi, previously known from a few individuals from the type locality in the SW Gulf of California and now collected in the SE Gulf of California, Mexico, is used to complete its description, particularly the morphology of thoracopods of both sexes. The new material matches the original description provided for this species, with very few, small variations. Variation of setal armature of telson posterior margin and uropodal endopod is provided based on the observation of 20 males, 20 females, and 20 ovigerous females of different size and the additional morphological data now available are compared with the two other species of the genus Cubanomysis. Additional information is provided regarding the size of the species, its coloration, habitat and reproductive aspects.

Australian Opilonini (Coleoptera: Cleridae: Clerinae) part II: description and taxonomy of Megaxestodes gen. nov. and introduction to an informal ‘Zenithicola-complex’ including taxonomic notes and new synonymies

JUSTIN S. BARTLETT — 2026-06-09

The species constitution and taxonomy of Zenithicola Spinola, 1841 is reviewed, resulting in description of Megaxestodes gen. nov.. for Megaxestodes crassus (Newman, 1840) comb. nov., Megaxestodes funestus (Chevrolat, 1874) comb. nov. and Megaxestodes socius (Chevrolat, 1874) stat. rev., comb. nov., the latter removed from synonymy with M. funestus. Zenithicola cribricollis Pic, 1941 syn. nov. is synonymised with Megaxestodes socius (Chevrolat, 1874) comb. nov., and Clerus obesus White, 1846 syn. nov. removed from synonymy with Megaxestodes crassus and newly synonymised with Megaxestodes funestus. The genera Balcus Sharp, 1877, Mimolesterus Gerstmeier, 1991, Phlogistomorpha Hintz, 1908, Phlogistus Gorham, 1876 and Zenithicola Spinola, 1841 are discussed within the context of a newly defined informal group, the Zenithicola-complex. The taxonomy of several Zenithicola-complex species is revised. New synonymies and combinations include: Aulicus dives Blackburn, 1900 syn. nov. = Phlogistomorpha mastersii (Macleay, 1872); Phlogistus affinis (Gorham, 1876) stat. rev., comb. nov. (removed from synonymy with Phlogistus smaragdinus (Gorham, 1876)); Aulicus corallipes Chevrolat, 1874 syn. nov. = Phlogistus castanipes (Westwood, 1849) stat. rev., comb. nov. (removed from synonymy with Phlogistus instabilis (Newman)); Phlogistus corporaali Pic, 1941 syn. nov. = Phlogistus sculptus (Macleay, 1872); Phlogistus rufimembris Pic, 1941 syn. nov. = Phlogistus niger Hintz, 1908; Phlogistus speciosus Hintz, 1908 syn. nov. = Phlogistus schulzi Hintz, 1908; Aulicus mundus Blackburn, 1900 syn. nov. = Phlogistus viridissimus (Pascoe, 1860) stat. rev., comb. nov. (removed from synonymy with Phlogistus delicatulus (Boheman, 1859)). Xylotretus scrobiculatus Spinola, 1845, hitherto assigned to Zenithicola, is classified as Cleridae incertae sedis. Lectotypes are designated for Clerus obesus White, 1846, Phlogistus remotus Schenkling, 1915, Zenithicola cribricollis Pic, 1941, Zenithicola funestus Chevrolat, 1874 and Zenithicola socius Chevrolat, 1874. Figures of Megaxestodes species and other taxa under discussion are presented.

**Redescription of Gyponana germari (Stål) (Hemiptera: Cicadellidae), a Neotropical species recently adventive in the continental USA, with notes on its distribution, ecology, and behavior**

SOLOMON V. HENDRIX — 2026-06-09

The Mexican leafhopper Gyponana germari (Stål), previously reported in Hawaii, Mexico, and Central America, has recently been documented in the continental United States from Florida. The species is here reported for the first time from Belize, El Salvador, and Honduras. Diagnosis of G. germari and its distinction from other similar Gyponana spp. in Florida are discussed. DNA sequence data for the loci 18S and 28S are reported for the first time, and the species is redescribed in detail. Potential interactions based on community science observations between G. germari and ants are briefly noted. The richness of the leafhopper genus Gyponana and adventive leafhoppers in Florida is briefly reviewed.

Erschoffia Swinhoe, 1900, a forgotten genus reinstated from synonymy with Aspitates Treitschke, 1825 (Lepidoptera: Geometridae, Ennominae)

IGOR KOSTJUK — 2026-06-09

The genus Erschoffia Swinhoe, 1900 is reinstated from synonymy with Aspitates Treitschke, 1825 to accomodate four ennomine species characterized by asymmetrical male genitalia. Erschoffia remmi, sp. nov. is described from the Western Tien Shan. E. acuminaria (Eversmann, 1851) exhibits a disjunct distribution between the highlands of Middle Asia and the Caucasus. The distribution and taxonomic significance of four key diagnostic characters of Erschoffia within Ennominae are analyzed.

**Taxonomic notes on the genus Antimimistis Turner (Lepidoptera: Geometridae: Larentiinae), with description of two new species from Southeast Asia**

OLGA SCHMIDT — 2026-06-09

The present study deals with the morphological characteristics of the Indo-Australian geometrid moth genus Antimimistis Turner, 1922 sensu Holloway, 1997. The genus currently comprises four species. The moths are small, the upper side of the wings is usually dark brown, with irregular whitish lines, often with a pale central spot at the termen. Two species, A. circumsubteracta, sp. nov. and A. malukensis, sp. nov., are described from Southeast Asia as new. The diagnostic attributes of the species are explained and illustrated herein.

Taxonomic contribution to the enigmatic ladybird beetle tribe Limnichopharini Miyatake, 1994 (Coleoptera: Coccinellidae), with description of the third species of Limnichopharus Miyatake, 1994 from Thailand

RYŌTA SEKI — 2026-06-09

A new species of the extremely rarely collected genus Limnichopharus Miyatake, 1994, of the monotypic, ladybird beetle tribe Limnichopharini Miyatake, 1994 (subfamily Coccinellinae), Limnichopharus khaoyaiensis sp. nov., is described from Thailand, illustrated, and compared with the other congeneric species L. hainanensis Miyatake and L. borneensis Miyatake. This represents the third known species in the genus and the tribe. Male genitalia are illustrated and described for the first time for Limnichopharini. The systematic position of the tribe is briefly discussed.

**On the synonymy of Trimeresurus liqibini Liang, Ding, Wu, Yang & Chen, 2026 with Trimeresurus loong Xu, Deng, Zhang, Nguyen, Poyarkov, Vogel & Peng, 2025 (Squamata: Viperidae)**

TAN VAN NGUYEN — 2026-06-09