Celante, G.L., Carvalho, A.G., Jahyny, B.J.B., Araújo, K.B.S. & Santos, P.B.D. (2024) A new species of Athyreus MacLeay, 1819 (Coleoptera: Geotrupidae: Bolboceratinae: Athyreini) from northeastern Brazil. Zootaxa, 5512 (3), 389–401.