Aguila, R.N., Genaro, J.A., Peréz-Asso, A.R. & Hausmann, A. (2017)

A new species of the hysius species-group of Calisto Hübner (Lepidoptera, Nymphalidae, Satyrinae) and insights into the status of different populations currently attributed to C. grannus Bates

. Zootaxa, 4317 (1), 1–44.