Maddison DR, Arnold AE. A review of the Bembidion (Odontium) aenulum subgroup (Coleoptera: Carabidae), with description of a new species. Zootaxa [Internet]. 2009 Aug. 31 [cited 2025 Sep. 6];2214(1):45–61. Available from: https://www.mapress.com/zt/article/view/zootaxa.2214.1.3