Vaz-De-Mello, Fernando Z., W. D. Edmonds, Federico C. Ocampo, and Paul Schoolmeesters. “A Multilingual Key to the Genera and Subgenera of the Subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae)”. Zootaxa 2854, no. 1 (April 29, 2011): 1–73. Accessed May 1, 2025. https://www.mapress.com/zt/article/view/zootaxa.2854.1.1.