Celante, Gabriel Luiz, Adhan Gabriel Carvalho, Benoit Jean Bernard Jahyny, Kaylla Brisley Silva Araújo, and Paula Batista Dos Santos. “A New Species of Athyreus MacLeay, 1819 (Coleoptera: Geotrupidae: Bolboceratinae: Athyreini) from Northeastern Brazil”. Zootaxa 5512, no. 3 (September 20, 2024): 389–401. Accessed May 1, 2025. https://www.mapress.com/zt/article/view/zootaxa.5512.3.4.