Mondaca, José. “A Checklist of the Scarabaeoidea (Coleoptera) of Chile With Exemplar Live-Photographed”. Zootaxa 5285, no. 2 (May 16, 2023): 201–251. Accessed May 1, 2025. https://www.mapress.com/zt/article/view/zootaxa.5285.2.1.