Erwin, Terry L. “The Beetle Family Carabidae of Costa Rica and Panam : Descriptions of Four New Genera and Six New Species With Notes on Their Way of Life (Insecta: Coleoptera)”. Zootaxa 537, no. 1 (June 14, 2004): 1–18. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.537.1.1.