Brown, Jason L., and Evan Twomey. “Complicated Histories: Three New Species of Poison Frogs of the Genus Ameerega (Anura: Dendrobatidae) from North-Central Peru”. Zootaxa 2049, no. 1 (March 23, 2009): 1–38. Accessed May 2, 2025. https://www.mapress.com/zt/article/view/zootaxa.2049.1.1.