Joly, Luis J. “Review of the Species in the Cyclocephala Bicolor Laporte Species Group (Coleoptera: Scarabaeidae: Dynastinae)”. Zootaxa 2048, no. 1 (March 23, 2009): 47–64. Accessed May 2, 2025. https://www.mapress.com/zt/article/view/zootaxa.2048.1.3.