Maldaner, Maria E., Rafael V. Nunes, and Fernando Z. Vaz-De-Mello. “Taxonomic Revision of the Dichotomius speciosus (Waterhouse, 1891) Species Group (Coleoptera: Scarabaeidae: Scarabaeinae)”. Zootaxa 3986, no. 5 (July 21, 2015): 549–560. Accessed May 2, 2025. https://www.mapress.com/zt/article/view/zootaxa.3986.5.2.