Kamiński, Marcin Jan. “A Taxonomic Revision of the Quadrideres simplicipes Species-Group (Coleoptera: Tenebrionidae: Pedinini), Including Description of a New Species from Tanzania”. Zootaxa 3986, no. 2 (July 16, 2015): 217–226. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.3986.2.5.