Heinicke, Matthew P., César L. Barrio-Amorós, and S. Blair Hedges. “Molecular and Morphological Data Support Recognition of a New Genus of New World Direct-Developing Frog (Anura: Terrarana) from an under-Sampled Region of South America”. Zootaxa 3986, no. 2 (July 16, 2015): 151–172. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.3986.2.1.