Ota, Rafaela P., Flávio C. T. Lima, and Carla S. Pavanelli. “A New Species of Hemigrammus Gill, 1858 (Characiformes: Characidae) from the Central and Western Amazon and Rio Paraná-Paraguai Basins”. Zootaxa 3948, no. 2 (April 21, 2015): 218–232. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.3948.2.4.