Hardy, Martin. “Description of a New Species of Bothynus Hope from Argentina and Bolivia (Coleoptera: Scarabaeidae: Dynastinae)”. Zootaxa 4362, no. 1 (December 4, 2017): 141–145. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.4362.1.9.