Reid, C. A. M., and M. Beatson. “Revision of the Stag Beetle Genus Ryssonotus MacLeay (Coleoptera: Lucanidae), With Descriptions of a New Genus and Three New Species”. Zootaxa 4150, no. 1 (August 10, 2016): 1–39. Accessed May 2, 2025. https://www.mapress.com/zt/article/view/zootaxa.4150.1.1.