Sekiya, Kunio, Ikuo Miura, and Mitsuaki Ogata. “A New Frog Species of the Genus Rugosa From Sado Island, Japan (Anura, Ranidae)”. Zootaxa 3575, no. 1 (December 7, 2012): 49–62. Accessed May 2, 2025. https://www.mapress.com/zt/article/view/zootaxa.3575.1.3.