Bilton, David T, and Philip D Perkins. “Two New Species of Parhydraenini from South Africa (Coleoptera: Hydraenidae)”. Zootaxa 3342, no. 1 (June 12, 2012): 51–59. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.3342.1.3.