Perkins, Philip D. “New Records and Description of Fifty-Four New Species of Aquatic Beetles in the Genus Hydraena Kugelann from South America (Coleoptera: Hydraenidae)”. Zootaxa 3074, no. 1 (October 28, 2011): 1–198. Accessed May 1, 2025. https://www.mapress.com/zt/article/view/zootaxa.3074.1.1.