Barrio-Amorós, César L., Ramón Rivero, and Juan C. Santos. “A New Striking Dendrobatid Frog (Dendrobatidae: Aromobatinae, Aromobates) from the Venezuelan Andes”. Zootaxa 3063, no. 1 (October 20, 2011): 39–52. Accessed May 3, 2025. https://www.mapress.com/zt/article/view/zootaxa.3063.1.3.