Kok, P. J. R., R. D. Macculloch, A. Lathrop, B. Willaert, and F. Bossuyt. “A New Species of Anomaloglossus (Anura: Aromobatidae) from the Pakaraima Mountains of Guyana”. Zootaxa, vol. 2660, no. 1, Oct. 2010, pp. 18–32, doi:10.11646/zootaxa.2660.1.2.