Landolt, P. J., J. M. Sierra, T. R. Unruh, and R. S. Zack. “A New Species of Vespula, and First Record of Vespa Crabro L. (Hymenoptera: Vespidae) from Guatemala, Central America”. Zootaxa, vol. 2629, no. 1, Sept. 2010, pp. 61–68, doi:10.11646/zootaxa.2629.1.4.