Pyron, R. A., and D. A. Beamer. “Systematic Revision of the Spotted and Northern Dusky Salamanders (Plethodontidae: Desmognathus Conanti and D. fuscus), With Six New Species from the Eastern United States”. Zootaxa, vol. 5311, no. 4, July 2023, pp. 451-04, doi:10.11646/zootaxa.5311.4.1.