Kamal, N. S. S., H. H. Tan, and C. K. C. Ng. “Betta nuluhon, a New Species of Fighting Fish from Western Sabah, Malaysia (Teleostei: Osphronemidae)”. Zootaxa, vol. 4819, no. 1, July 2020, pp. 187–194, doi:10.11646/zootaxa.4819.1.11.