Rahimian, H., S. Shafiei, N. R. Pouyani, and E. R. Pouyani. “Phylogenetic Relationships of the Gray-Toad Agama, Phrynocephalus scutellatus (Olivier, 1807), Species Complex from Iran”. Zootaxa, vol. 3990, no. 3, July 2015, pp. 369–380, doi:10.11646/zootaxa.3990.3.3.