Dehling, J. M., and M. Matsui. “A New Species of Leptolalax (Anura: Megophryidae) from Gunung Mulu National Park, Sarawak, East Malaysia (Borneo)”. Zootaxa, vol. 3670, no. 1, June 2013, pp. 33–44, doi:10.11646/zootaxa.3670.1.2.