Sathiandran, N., and T. K. Sabu. “New Species, New Synonym, and Redescription of Onthophagus (Coleoptera: Scarabaeidae: Scarabaeinae) from the Western Ghats, India”. Zootaxa, vol. 3526, no. 1, Oct. 2012, pp. 53–58, doi:10.11646/zootaxa.3526.1.5.