Plisko, J. D. “An Annotated Checklist of the South African Acanthodrilidae (Oligochaeta: Acanthodrilidae: Acanthodrilinae, Benhamiinae)”. Zootaxa, vol. 3458, no. 1, Sept. 2012, pp. 4–58, doi:10.11646/zootaxa.3458.1.2.