J. M. S. Costas, L. L. Moreda, and C. G. Luque, “A new cave-dwelling species of Quaestus Schaufuss, 1861 from the Montes de Pas, Northern Spain (Coleoptera: Leiodidae: Leptodirini)”, Zootaxa, vol. 2484, no. 1, pp. 25–34, May 2010.