Y. Liu, H. Liang, D. H. Kavanaugh, and M. Yang, “Key to species of the subgenus Chlaenioctenus (Coleoptera: Carabidae: Chlaeniini: Chlaenius), with description of two new species”, Zootaxa, vol. 2397, no. 1, pp. 15–28, Mar. 2010.