I. V. Enushchenko and V. B. Semenov, “A review of the genus Gyrophaena Mannerheim 1830 (Coleoptera: Staphylinidae: Aleocharinae: Gyrophaenina) of the Caucasus and adjacent territories”, Zootaxa, vol. 4126, no. 3, pp. 301–337, Jun. 2016.