Pombal Jr., J. P., Siqueira, C. C., Dorigo, T. A., Vrcibradic, D. and Rocha, C. F. D. (2008) “A third species of the rare frog genus Holoaden (Terrarana, Strabomantidae) from a montane rainforest area of southeastern Brazil”, Zootaxa. Auckland, New Zealand, 1938(1), pp. 61–68. doi: 10.11646/zootaxa.1938.1.4.